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The song thrush ( Turdus philomelos) is a thrush that breeds across the West Palearctic. It has brown upper-parts and black-spotted cream or buff underparts and has four recognised subspecies. Its distinctive Birdsong, which has repeated musical phrases, has frequently been referred to in poetry.
The song thrush breeds in forests, gardens and parks, and is partially bird migration with many birds wintering in southern Europe, North Africa and the Middle East; it has also been introduced into New Zealand and Australia. Although it is not threatened globally, there have been serious population declines in parts of Europe, possibly due to changes in farming practices.
The song thrush builds a neat mud-lined cup nest in a bush or tree and lays four to five dark-spotted blue bird egg. It is omnivorous and has the habit of using a favourite stone as an "anvil" on which to break open the shells of land snail. Like other perching birds (), it is affected by external and internal and is vulnerable to predation by and birds of prey.
The song thrush has four subspecies, with the nominate subspecies, T. p. philomelos, covering the majority of the species' range. T. p. hebridensis, described by United Kingdom ornithologist William Eagle Clarke in 1913, is a mainly sedentism (non-migratory) form found in the Outer Hebrides and Isle of Skye in Scotland, and western Ireland. It is the darkest subspecies, with a dark brown back, greyish rump, pale buff background colour to the underparts and grey-tinged flanks.
T. p. clarkei, described by German zoologist Ernst Hartert in 1909, and named after William Eagle Clarke, breeds in the rest of Great Britain and Ireland and on mainland Europe in France, Belgium, the Netherlands and possibly somewhat further east. It has brown upperparts which are warmer in tone than those of the nominate form, an olive-tinged rump and rich yellow background colour to the underparts. It is a partial migrant with some birds wintering in southern France and Iberia. This form intergrades with the nominate subspecies in central Europe, and with T. p. hebridensis in the Inner Hebrides and western Scotland, and in these areas birds show intermediate characteristics. Finally, T. p. nataliae, described by the Russian Sergei Buturlin in 1929, occurs in the east of the range in Siberia east of the Ural Mountains.
The most similar European thrush species is the redwing ( T. iliacus), but that bird has a strong white supercilium, red flanks, and shows a red underwing in flight. The mistle thrush ( T. viscivorus) is much larger and has white tail corners, and the Chinese thrush ( T. mupinensis), although much more similar in plumage, has black face markings and does not overlap in range.
The song thrush has a short, sharp tsip call, replaced on migration by a thin high seep, similar to the redwing's call but shorter. The alarm call is a chook-chook becoming shorter and more strident with increasing danger. The male's song, given from trees, rooftops or other elevated perches, is a loud clear run of musical phrases, repeated two to four times, filip filip filip codidio codidio quitquiquit tittit tittit tereret tereret tereret, and interspersed with grating notes and mimicry. It is given mainly from February to June by the Outer Hebridean race, but from November to July by the more widespread subspecies. For its weight, this species has one of the loudest bird calls.
An individual male may have a repertoire of more than 100 phrases, many copied from its parents and neighbouring birds. Mimicry may include the imitation of man-made items like telephones, and the song thrush will also repeat the calls of captive birds, including exotics such as the white-faced whistling duck.
In Great Britain song thrushes are commonly found where there are trees and bushes. Such areas include parks, gardens, coniferous and deciduous woodland and hedgerows.
Birds of the subspecies T. p. clarkei were introduced to New Zealand and Australia by acclimatisation societies between 1860 and 1880, apparently for purely sentimental reasons. In New Zealand, where it was introduced on both the main islands, the song thrush quickly established itself and spread to surrounding islands such as the Kermadec Islands, Chatham Islands and Auckland Islands. Although it is common and widespread in New Zealand, in Australia only a small population survives around Melbourne. In New Zealand, there appears to be a limited detrimental effect on some invertebrates due to predation by introduced bird species, and the song thrush also damages commercial fruit crops in that country. As an introduced species it has no legal protection in New Zealand, and can be killed at any time.
The song thrush typically nests in forest with good undergrowth and nearby more open areas, and in western Europe also uses gardens and parks. It breeds up to the tree-line, reaching in Switzerland. The island subspecies T. p. hebridensis breeds in more open country, including heathland, and in the east of the song thrush's Eurasian range, the nominate subspecies is restricted to the edge of the dense conifer forests.
In intensively farmed areas where agricultural practices appear to have made cropped land unsuitable, gardens are an important breeding habitat. In one English study, only 3.5% of territories were found in farmland, whereas gardens held 71.5% of the territories, despite that habitat making up only 2% of the total area. The remaining nests were in woodlands (1% of total area).
The winter habitat is similar to that used for breeding, except that high ground and other exposed localities are avoided; however, the island subspecies T. p. hebridensis will frequent the seashore in winter.
This is a monogamy territorial species, and in areas where it is fully migratory, the male re-establishes its breeding territory and starts singing as soon as he returns. In the milder areas where some birds stay year round, the resident male remains in his breeding territory, singing intermittently, but the female may establish a separate individual wintering range until pair formation begins in the early spring.
During migration, the song thrush travels mainly at night with a strong and direct flight action. It flies in loose flocks which cross the sea on a broad front rather than concentrating at short crossings (as occurs in the migration of large soaring birds), and calls frequently to maintain contact. Migration may start as early as late August in the most easterly and northerly parts of the range, but the majority of birds, with shorter distances to cover, head south from September to mid-December. However, hard weather may force further movement. Return migration varies between mid-February around the Mediterranean to May in northern Sweden and central Siberia.
Adult birds may be killed by , and sparrowhawks, and eggs and nestlings are taken by European magpie, Eurasian jay, and, where present, grey squirrels. As with other passerine birds, parasites are common, and include endoparasites, such as the nematode Splendidofilaria (Avifilaria) mavis whose specific name mavis derives from this thrush. A Russian study of blood parasites showed that all the fieldfares, redwings and song thrushes sampled carried haematozoans, particularly Haemoproteus and Trypanosoma. Ixodes ticks are also common, and can carry pathogens, including tick-borne encephalitis in forested areas of central and eastern Europe and Russia, and, more widely, Borrelia bacteria. Some species of Borrelia cause Lyme disease, and ground-feeding birds like the song thrush may act as a reservoir for the disease.
are an especially important food item when drought or hard weather makes it hard to find other food. The thrush often uses a favorite stone as an "anvil" on which to break the shell of the snail before extracting the soft body and invariably wiping it on the ground before consumption. Young birds initially flick objects and attempt to play with them until they learn to use anvils as tools to smash snails. The nestlings are mainly fed on animal food such as worms, , snails and insect .
The grove snail ( Cepaea nemoralis) is regularly eaten by the song thrush, and its polymorphic shell patterns have been suggested as evolutionary responses to reduce predation; however, song thrushes may not be the only selective force involved.
In the western Palaearctic, there is evidence of population decline, but at a level below the threshold required for global conservation concern (i.e., a reduction in numbers of more than 30% in ten years or three generations) and the IUCN Red List categorises this species as of "Least Concern". In Great Britain and the Netherlands, there has been a more than 50% decline in population, and the song thrush is included in regional Red Lists. The decreases are greatest in farmlands (73% since the mid-1970s) and believed to be due to changes in agricultural practices in recent decades. The precise reasons for the decline are not known but may be related to the loss of hedgerows, a move to sowing crops in autumn rather than spring, and possibly the increased use of pesticides. These changes may have reduced the availability of food and of nest sites. In gardens, the use of poison bait to control slugs and snails may pose a threat. In urban areas, some thrushes are road kill while using the hard surface of roads to smash snails.
That's the wise thrush; he sings each song twice over,The song also inspired the nineteenth-century British writer Thomas Hardy, who spoke in Darkling Thrush of the bird's "full-hearted song evensong/Of joy illimited", but twentieth-century British poet Ted Hughes in Thrushes concentrated on its hunting prowess: "Nothing but bounce and/stab/and a ravening second". Nineteenth-century Welsh poet Edward Thomas wrote 15 poems concerning blackbirds or thrushes, including The Thrush:
Lest you should think he never could recapture
The first fine careless rapture!
I hear the thrush, and I see
Him alone at the end of the lane
Near the bare poplar's tip,
Singing continuously.
In The Tables Turned, Romanticism poet William Wordsworth references the song thrush, writing
Hark, how blithe the throstle sings
And he is no mean preacher
Come forth into the light of things
Let Nature be your teacher
The song thrush is the emblem of West Bromwich Albion Football Club, chosen because the public house in which the team used to change kept a pet thrush in a cage. It also gave rise to Albion's early nickname, The Throstles. A few English pubs and hotels share the name Throstles Nest.
In Spain, this species is normally caught as it migrates through the country, often using birdlime which, although banned by the European Union, is still tolerated and permitted in the Valencian Community. In 2003 and 2004 the European Union tried, but failed, to stop this practice in the Valencian region.
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