Product Code Database
Example Keywords: mobile -office $61-103
barcode-scavenger
   » » Wiki: Aurochs
Tag Wiki 'Aurochs'.
Tag

The aurochs ( Bos primigenius; or ; pl.: aurochs or aurochsen) is an species of , considered to be the wild ancestor of modern domestic . With a shoulder height of up to in bulls and in cows, it was one of the largest in the ; it had massive elongated and broad horns that reached in length.

The aurochs was part of the Pleistocene megafauna. It probably evolved in Asia and migrated west and north during warm periods. The oldest-known aurochs fossils date to the Middle Pleistocene. The species had an expansive range spanning from Western Europe and North Africa to the Indian subcontinent and East Asia. The distribution of the aurochs progressively contracted during the Holocene due to and hunting, with the last known individual dying in the Jaktorów forest in Poland in 1627.

There is a long history of interaction between aurochs and humans, including archaic hominins like . The aurochs is depicted in , , and figurines. It symbolised power, sexual potency and prowess in religions of the ancient Near East. Its horns were used in , as and .

Two aurochs events occurred during the Neolithic Revolution. One gave rise to the domestic ( Bos taurus) in the in the that was introduced to Europe via the and the coast of the Mediterranean Sea. Hybridisation between aurochs and early domestic cattle occurred during the early Holocene. Domestication of the led to the cattle ( Bos indicus) that hybridised with early taurine cattle in the about 4,000 years ago. Some modern cattle breeds exhibit features reminiscent of the aurochs, such as the dark colour and light eel stripe along the back of bulls, the lighter colour of cows, or an aurochs-like horn shape.


Etymology
Both "aur" and "ur" are Germanic or words meaning "wild ox".
(1983). 9780517414255, Greenwich House.
In Old High German, this word was compounded with ohso ('ox') to ūrohso, which became the early modern Aurochs. The word "urus" was used for wild ox from the onwards.

The use of the plural form aurochsen in English is a direct parallel of the German plural Ochsen and recreates the same distinction by analogy as English singular ox and plural oxen, although aurochs may stand for both the singular and the plural term; both are attested.

(2025). 9781108437738, Cambridge University Press.
(2025). 9783319695785, Palgrave Macmillan.


Taxonomy and evolution
The Bos taurus was introduced by in 1758 for cattle in Poland. The scientific name Bos primigenius was proposed for the aurochs by Ludwig Heinrich Bojanus who described the skeletal differences between the aurochs and domestic cattle in 1825, published in 1827. The name Bos namadicus was used by in 1859 for cattle found in deposits. Bos primigenius mauritanicus was coined by in 1881 who described fossils found in deposits near west of Constantine, Algeria.

In 2003, the International Commission on Zoological Nomenclature placed Bos primigenius on the Official List of Specific Names in Zoology and thereby recognized the validity of this name for a wild species.


Subspecies
Three aurochs have traditionally been recognised to have existed in historical times:
  • The Eurasian aurochs ( B. p. primigenius) was part of the Pleistocene megafauna in and survived until the 17th century in .
  • The ( B. p. namadicus) lived on the Indian subcontinent.
    (2025). 9780309139861
  • The North African aurochs ( B. p. mauritanicus) lived north of the . This subspecies has also been called B. p. opisthonomus.
Chinese geneticists published mitochondrial DNA evidence supporting that Eurasian aurochs populations from were genetically isolated for large stretches of the , and as a result distinctive enough to be considered a separate subspecies, the East Asian aurochs ( B. p. sinensis), even if this was not morphologically distinct.

At least two of aurochs developed on Mediterranean islands as a result of sea level changes during the Pleistocene:

  • B. p. siciliae on the Italian island of .
  • B. p. thrinacius on the Greek island of .


Evolution
Calibrations using of 16 Bovidae species indicate that the tribe evolved about . The and are estimated to have genetically diverged from the Bovini about . The following shows the relationships of the aurochs based on analysis of and mitochondrial genomes in the Bovini tribe:

The cold caused an extension of open , which enabled the of large grazers.

(2025). 9789546422354, Pensoft Publishers.
The origin of the aurochs is unclear, with authors suggesting either an African or Asian origin for the species. is considered to be a possible ancestor of the aurochs, of which a fossil skull was excavated in the in India that dates to the Early Pleistocene about .

An aurochs skull excavated in Tunisia's from early Middle Pleistocene dating about is the oldest well-dated fossil specimen to date. The authors of the study proposed that Bos might have evolved in Africa and migrated to Eurasia during the Middle Pleistocene. Middle Pleistocene aurochs fossils were also excavated in a erg in the .

Fossils of the Indian subspecies ( Bos primigenius namadicus) were excavated in in dating to the Middle Pleistocene. Remains of aurochs are common in Late Pleistocene sites across the Indian subcontinent.

The earliest fossils in Europe date to the Middle Pleistocene. One site widely historically suggested to represent the first appearance of aurochs in Europe was the Notarchirico site in southern Italy, dating around 600,000 years ago, however a 2024 re-examination of the site found that presence of aurochs at the locality was unsupported, with the oldest records of aurochs now placed at the Ponte Molle site in central Italy, dating to around 550–450,000 years ago. Aurochs were present in Britain by Marine Isotope Stage 11 ~400,000 years ago.

The earliest remains aurochs in East Asia are uncertain, but may date to the late Middle Pleistocene.

Late Pleistocene aurochs fossils were found in Affad 23 in Sudan dating to 50,000 years ago when the climate in this region was more humid than during the African humid period.

Following the most recent , the range of the aurochs expanded into Denmark and southern Sweden at the beginning of the Holocene, around 12–11,000 years ago.


Description
According to a 16th-century description by Sigismund von Herberstein, the aurochs was pitch-black with a grey streak along the back; his wood carving made in 1556 was based on a culled aurochs, which he had received in . In 1827, Charles Hamilton Smith published an image of an aurochs that was based on an that he had purchased from a merchant in , which is thought to have been made in the early 16th century. This painting is thought to have shown an aurochs, although some authors suggested it may have shown a hybrid between an aurochs and domestic cattle, or a Polish steer. Contemporary reconstructions of the aurochs are based on and the information derived from contemporaneous artistic depictions and historic descriptions of the animal.


Coat colour
Remains of aurochs hair were not known until the early 1980s. Depictions show that the North African aurochs may have had a light saddle marking on its back.
(2025). 9783000267642, Lipp Graphische Betriebe.
Calves were probably born with a chestnut colour, and young bulls changed to black with a white running down the spine, while cows retained a reddish-brown colour. Both sexes had a light-coloured muzzle, but evidence for variation in coat colour does not exist. Egyptian grave paintings show cattle with a reddish-brown coat colour in both sexes, with a light saddle, but the horn shape of these suggest that they may depict domesticated cattle. Many primitive cattle breeds, particularly those from Southern Europe, display similar coat colours to the aurochs, including the black colour in bulls with a light eel stripe, a pale mouth, and similar sexual dimorphism in colour. A feature often attributed to the aurochs is blond forehead hairs. According to historical descriptions of the aurochs, it had long and curly forehead hair, but none mentions a certain colour. Although the colour is present in a variety of primitive cattle breeds, it is probably a discolouration that appeared after domestication.


Body shape
The proportions and body shape of the aurochs were strikingly different from many modern cattle breeds. For example, the legs were considerably longer and more slender, resulting in a shoulder height that nearly equalled the trunk length. The skull, carrying the large horns, was substantially larger and more elongated than in most cattle breeds. As in other wild bovines, the body shape of the aurochs was athletic, and especially in bulls, showed a strongly expressed neck and shoulder musculature. Therefore, the fore hand was larger than the rear, similar to the wisent, but unlike many domesticated cattle. Even in carrying cows, the udder was small and hardly visible from the side; this feature is equal to that of other wild bovines.


Size
The aurochs was one of the largest in Europe. The size of an aurochs appears to have varied by region, with larger specimens in northern Europe than farther south. Aurochs in Denmark and Germany ranged in height at the shoulders between in bulls and in cows, while aurochs bulls in reached .

The African aurochs was similar in size to the European aurochs in the Pleistocene, but declined in size during the transition to the Holocene; it may have also varied in size geographically.

The body mass of aurochs appears to have shown some variability. Some individuals reached around , whereas those from the late Middle Pleistocene are estimated to have weighed up to . The aurochs exhibited considerable sexual dimorphism in the size of males and females.


Horns
The horns were massive, reaching in length and between in diameter. Its horns grew from the skull at a 60-degree angle to the muzzle facing forwards and were curved in three directions, namely upwards and outwards at the base, then swinging forwards and inwards, then inwards and upwards. The curvature of bull horns was more strongly expressed than horns of cows. The basal circumference of reached in the largest Chinese specimen and in a French specimen. Some cattle breeds still show horn shapes similar to that of the aurochs, such as the Spanish fighting bull, and occasionally also individuals of derived breeds.


Genetics
A well-preserved aurochs bone yielded sufficient mitochondrial DNA for a sequence analysis in 2010, which showed that its genome consists of 16,338 . Further studies using the aurochs whole genome sequence have identified candidate microRNA-regulated domestication genes. A comprehensive sequence analysis of Late Pleistocene and Holocene aurochs published in 2024 suggested that Indian aurochs (represented by modern zebu cattle) were the most genetically divergent aurochs population, having diverged from other aurochs around 300–166,000 years ago, with other aurochs populations spanning Europe and the Middle East to East Asia sharing much more recent common ancestry within the last 100,000 years. Late Pleistocene European aurochs were found to have a small (~3%) ancestry component from a divergent lineage that split prior to the divergence of Indian and other aurochs, suggested to be residual from earlier European aurochs populations. Towards the end of the Late Pleistocene, European aurochs experienced considerable gene flow from Middle Eastern aurochs. European Holocene aurochs primarily descend from those that were present in the Iberian Peninsula during the Last Glacial Maximum, with the Holocene also seeing mixing between previously isolated aurochs populations.


Distribution and habitat
The aurochs was widely distributed in , , and throughout Europe to the Pontic–Caspian steppe, and in the west and to the Gulf of Finland and in the north.

Fossil horns attributed to the aurochs were found in deposits at an elevation of on the eastern margin of the close to the in Zoigê County that date to about 26,620±600 years BP. Most fossils in were found in plains below in , Yushu, Jilin, northeastern , , near , in province, and in province, in and in provinces. in aurochs fossils found in indicate that the aurochs survived in the region until at least 5,000 years BP. Fossils were also excavated on ,

(2025). 9788968113697, Magnolia Press.
and in the Japanese archipelago.

During warm periods the aurochs was widespread across Europe, but during retreated into southern refugia in the Iberian, Italian and peninsulas.

Landscapes in Europe probably consisted of dense forests throughout much of the last few thousand years. The aurochs is likely to have used and along lakes. Analysis of specimens found in Britain suggests that aurochs preferred inhabiting low lying relatively flat landscapes. of mostly small found in fossiliferous with aurochs remains in China indicate that it preferred temperate grassy plains or bordering . It may have also lived in open grasslands. In the warm Atlantic period of the Holocene, it was restricted to remaining open country and forest margins, where competition with livestock and humans gradually increased leading to a successive decline of the aurochs.


Behaviour and ecology
Aurochs formed small herds mainly in winter, but typically lived singly or in smaller groups during the summer. If aurochs had social behaviour similar to their descendants, social status would have been gained through displays and fights, in which both cows and bulls engaged. Since it has a jaw, it has been suggested to have been a grazer, with a food selection very similar to domesticated cattle feeding on grass, twigs and . analysis of Holocene Danish aurochs teeth indicates that it changed from an abrasion-dominated grazer in the Danish to a mixed feeder in the Boreal, Atlantic and periods. and mesowear analysis of specimens from the Pleistocene of Britain has found these aurochs had mixed feeding to browsing diets, rather than being strict grazers.

season was in September, and calves were born in spring. Rutting bulls had violent fights, and evidence from the Jaktorów forest shows that they were fully capable of mortally wounding one another. In autumn, aurochs fed for the winter, gaining weight and possessing a shinier coat than during the rest of the year. Calves stayed with their mothers until they were strong enough to join and keep up with the herd on the feeding grounds. Aurochs calves would have been vulnerable to predation by ( Canis lupus) and ( Ursus arctos), while the immense size and strength of healthy adult aurochs meant they likely did not need to fear most predators. According to historical descriptions, the aurochs was swift despite its build, could be very aggressive if provoked, and was not generally fearful of humans. In Middle Pleistocene Europe, aurochs were likely predated upon by the "European jaguar" Panthera gombaszogensis and the ( Homotherium latidens), with evidence for the consumption of aurochs by ( Crocuta ( Crocuta) spelaea) having been found from Late Pleistocene Italy. The ( Panthera leo), ( Panthera tigris) and wolf are thought to have been the aurochs' main predators during the Holocene.

During periods in the Middle Pleistocene and early Late Pleistocene in Europe, the aurochs occurred alongside other large temperate adapted megafauna species, including the straight-tusked elephant ( Palaeoloxodon antiquus), Merck's rhinoceros ( Stephanorhinus kirchbergensis), the narrow-nosed rhinoceros, ( Stephanorhinus hemitoechus) and the ( Megaloceros giganteus).


Relationship with humans

In Asia
layers in on India's southern yielded aurochs bones with cut marks. An aurochs bone with cut marks induced with was found in a Middle Paleolithic layer at the Nesher Ramla Homo site in Israel; it was dated to Marine Isotope Stage 5 about 120,000 years ago. An archaeological excavation in Israel found traces of a feast held by the culture around 12,000 years BP, in which three aurochs were eaten. This appears to be an uncommon occurrence in the culture and was held in conjunction with the burial of an older woman, presumably of some social status. depicting aurochs in Gobustan Rock Art in Azerbaijan date to the Upper Paleolithic to periods. Aurochs bones and skulls found at the settlements of , Hallan Çemi and Çayönü indicate that people stored and shared food in the Pre-Pottery Neolithic B culture. Remains of an aurochs were also found in a in , Lebanon, dating to around 3,700 years BP; the aurochs was buried together with numerous animals, a few human bones and foods.

Seals dating to the Indus Valley civilisation found in and show an animal with curved horns like an aurochs.

(2025). 9789004168190, Brill.
Aurochs figurines were made by the in the .
(2025). 9781785707247, Oxbow Books.

The aurochs is denoted in the Akkadian words rīmu and rēmu, both used in the context of hunts by rulers such as Naram-Sin of Akkad, Tiglath-Pileser I and ; in Mesopotamia, it symbolised power and sexual potency, was an of the gods and , denoted prowess as an epithet of the king and the hero . Wild bulls are frequently referred to in as hunted by and to the god .

(2025). 9783868350876, Ugarit-Verlag.
An aurochs is depicted on 's , constructed in the 6th century BC.
(2025). 9780231537698, Columbia University Press.


In Africa
Petroglyphs depicting aurochs found in in the upper Nile valley were dated to the Late Pleistocene about 19–15,000 years BP using luminescence dating and are the oldest engravings found to date in Africa. Aurochs are part of hunting scenes in in a tomb at Thebes, Egypt dating to the 20th century BC, and in the of at dating to around 1175 BC. The latter is the youngest depiction of aurochs in art to date.


In Europe
Evidence has been found for the butchery of aurochs by archaic hominins in Europe during the Middle Palaeolithic, such as the Biache-Saint-Vaast site in northern France dating to around 240,000 years ago, where bones of aurochs have been found burnt by fire and with cut marks, thought to have been created by . At the late Middle Palaeolithic Cueva Des-Cubierta site in Spain, Neanderthals are proposed to have kept the skulls of aurochs as hunting trophies.

The aurochs is widely represented in Upper Paleolithic in the and caves in southern France dating to 36,000 and 21,000 years BP, respectively. Two Paleolithic in the depict an aurochs. Palaeolithic engravings showing aurochs were also found in the Grotta del Genovese on the Italian island of . Upper Paleolithic rock engravings and paintings depicting the aurochs were also found in caves on the Iberian Peninsula dating from the to the cultures.

(1999). 9783980583961, Neanderthal Museum.
(2025). 9781407303048, Archaeopress.
Aurochs bones with chop and cut marks were found at various hunting and butchering sites in France, Luxemburg, Germany, the Netherlands, England and Denmark. Aurochs bones were also found in Mesolithic settlements by the Narva and Emajõgi rivers in . Aurochs and human bones were uncovered from pits and at several Neolithic sites in England. A cup found in the Greek site of shows a hunting scene, in which people try to capture an aurochs. One of the bulls throws one hunter on the ground while attacking the second with its horns. The cup seems to date to . and hunted aurochs and used their huge horns as trophies, cups for wine, and offerings to the gods and heroes. The ox mentioned by Samus, Philippus of Thessalonica and as killed by Philip V of Macedon on the foothills of mountain , was actually an aurochs; Philip offered the horns, which were long, and the skin to a temple of .
(2025). 9780405084614, B. Blom.
The aurochs was described in 's Commentarii de Bello Gallico. Aurochs were occasionally captured and exhibited in shows in Roman amphitheatres such as the . Aurochs horns were often used by Romans as hunting horns.

In the , kills four aurochs.

(2025). 9783618661207, Deutscher Klassiker Verlag.
During the , aurochs horns were used as including the horn of the last bull; many aurochs horn sheaths are preserved today. The aurochs drinking horn at Corpus Christi College, Cambridge was engraved with the college's coat of arms in the 17th century. An aurochs head with a star between its horns and Christian iconographic elements represents the official coat of arms of Moldavia perpetuated for centuries.

Aurochs were hunted with arrows, nets and hunting dogs, and its hair on the forehead was cut from the living animal; belts were made out of this hair and believed to increase the of women. When the aurochs was slaughtered, the os cordis was extracted from the heart; this bone contributed to the mystique and magical powers that were attributed to it. In eastern Europe, the aurochs has left traces in expressions like "behaving like an aurochs" for a drunken person behaving badly, and "a bloke like an aurochs" for big and strong people.


Domestication
The earliest-known of the aurochs dates to the Neolithic Revolution in the , where cattle hunted and kept by Neolithic farmers gradually decreased in size between 9800 and 7500 BC. Aurochs bones found at and Göbekli Tepe are larger in size than cattle bones from later Neolithic settlements in northern like Dja'de el-Mughara and .
(2025). 9781842171219, Oxbow Books.
In sites of northern and western dating to the sixth millennium BC, cattle remains are also smaller but more frequent, indicating that domesticated cattle were imported during the from the central Fertile Crescent region. Results of genetic research indicate that the modern ( Bos taurus) arose from 80 aurochs in southeastern and northern Syria about 10,500 years ago. Taurine cattle spread into the and northern Italy along the River and the coast of the Mediterranean Sea. Hybridisation between male aurochs and early domestic cattle occurred in central Europe between 9500 and 1000 BC. Analyses of mitochondrial DNA sequences of Italian aurochs specimens dated to 17–7,000 years ago and 51 modern cattle breeds revealed some degree of of aurochs genes into south European cattle, indicating that female aurochs had contact with free-ranging domestic cattle. Cattle bones of various sizes found at a settlement in the Kutná Hora District provide further evidence for hybridisation of aurochs and domestic cattle between 3000 and 2800 BC in the region. Whole genome sequencing of a 6,750-year-old aurochs bone found in England was compared with genome sequence data of 81 cattle and single-nucleotide polymorphism data of 1,225 cattle. Results revealed that British and Irish cattle breeds share some with the aurochs specimen; early herders in Britain might have been responsible for the local from aurochs into the ancestors of British and Irish cattle. The cattle breed also exhibits sporadic introgression of female European aurochs into domestic cattle in the . Domestic cattle continued to diminish in both body and horn size until the Middle Ages.

Comparative analysis of single-nucleotide polymorphisms and shared revealed admixture between East Asian aurochs and introduced taurine cattle in , for example at . This suggested the incorporation of local aurochs into domestic cattle as far back as 4,000 years BP, either through spontaneous , or the capture of different aurochs groups to supplement domestic stocks. The same study detected derived alleles shared by aurochs and modern taurine cattle in East Asia, especially among breeds. Introgression with local aurochs could have facilitated rapid adaptation to new environments.

The Indian aurochs is thought to have been domesticated 10,000–8,000 years ago. Aurochs fossils found at the Neolithic site of in Pakistan are dated to around 8,000 years BP and represent some of the earliest evidence for its domestication on the Indian subcontinent. Female Indian aurochs contributed to the gene pool of ( Bos indicus) between 5,500 and 4,000 years BP during the expansion of in northern India. The zebu initially spread eastwards to . Hybridisation between zebu and early taurine cattle occurred in the after 4,000 years BP coinciding with the period during the 4.2-kiloyear event. The zebu was introduced to about 3,500–2,500 years ago, and reached in the 13th and 14th centuries.

A third domestication event thought to have occurred in Egypt's Western Desert is not supported by results of an analysis of genetic admixture, introgression and migration patterns of 3,196 domestic cattle representing 180 populations. However, the same study supported extensive hybridization between taurine cattle in Africa, arrived from the Near East after domestication, and local wild African aurochs prior to the entry of the zebu in Africa. The zebu was introduced through and started to spread comprehensively through in the last 1,400 years, along with cultural influences. Most modern breeds are hybridized to a variable extent with Indicine cattle, with introgression being most reduced in areas of West Africa where the is present.


Extinction
The Indian aurochs ( B. p. namadicus) became extinct sometime during the Holocene period, likely due to habitat loss caused by expanding and interbreeding with the domestic .
(2025). 9788178241401, Permanent Black.
The timing of extinction of aurochs in the Indian subcontinent is unclear, due to difficulty distinguishing aurochs remains from those of domestic cattle, with a 2021 review suggesting remains from Mehrgarh, Pakistan, dating to around 8,000 years ago "might constitute the only dated and reliably identified evidence" of Holocene Indian aurochs. The extinction probably predates the historical period, due to a lack of references to the aurochs in Indian texts.

A 2014 review suggested that the youngest remains of African aurochs ( B. p. mauritanicus) dated to around 6,000 years (BP), though some authors suggest that it may have survived until at least to the , as indicated by remains found in and in the .

In China, aurochs persisted until at least 3,600 BP.

The Eurasian aurochs ( B. p. primigenius) was present in southern Sweden during the Holocene climatic optimum until at least 7,800 years BP.

(2025). 9789402411058, Springer.
In Denmark, the first-known of the aurochs occurred after the sea level rise on the newly formed about 8,000–7,500 years BP, and the last documented aurochs lived in southern around 3,000 years BP. The latest-known aurochs fossil in dates to 3,245 years BP, and it was probably extinct by 3,000 years ago.

Excessive hunting began and continued until the aurochs was nearly extinct. The gradual extinction of the aurochs in was concurrent with the of large forest tracts between the 9th and 12th centuries.

By the 13th century, the aurochs existed only in small numbers in , and hunting it became a privilege of nobles and later royals. The population in was declining from at least the 9th century and was extinct in the 13th century.

Findings from records indicate that wild aurochs might have survived in northwestern until the 14th to 16th century, in until probably the early 17th century.

The last-known aurochs herd lived in a woodland in Poland's Jaktorów Forest. It decreased from around 50 individuals in the mid 16th century to four individuals by 1601. The last aurochs cow died in 1627 from natural causes.

A 2021 study argued that the aurochs possibly survived in northeastern until at least the 17th century. A horn-core excavated in 2020 in was identified as being from an aurochs; the archaeological layer in which it was found was dated to the second half of the 17th or first half of the 18th century, suggesting that aurochs may have survived in Bulgaria until that date.


Breeding of aurochs-like cattle
In the early 1920s, initiated a selective breeding program in attempting to the aurochs using several cattle breeds; the result is called . Herds of these cattle were released to Oostvaardersplassen, a in the Netherlands in the 1980s as aurochs surrogates for naturalistic grazing with the aim to restore prehistorical landscapes. Large numbers of them died of during the cold winters of 2005 and 2010, and the project of no interference ended in 2018.

Starting in 1996, Heck cattle were crossed with southern European cattle breeds such as , and to a lesser extent Spanish Fighting Bulls in the hope of creating a more aurochs-like animal. The resulting crossbreeds are called . Other breeding-back projects are the and the . However, approaches aiming at breeding an aurochs-like do not equate to an aurochs-like .


See also


External links

Page 1 of 1
1
Page 1 of 1
1

Account

Social:
Pages:  ..   .. 
Items:  .. 

Navigation

General: Atom Feed Atom Feed  .. 
Help:  ..   .. 
Category:  ..   .. 
Media:  ..   .. 
Posts:  ..   ..   .. 

Statistics

Page:  .. 
Summary:  .. 
1 Tags
10/10 Page Rank
5 Page Refs
1s Time