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Acer is a of and commonly known as maples. The is placed in the soapberry family .Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 and. http://www.mobot.org/MOBOT/research/APweb/. There are approximately 132 , most of which are native to , with a number also appearing in , northern , and . Only one species, , extends to the Southern Hemisphere.Gibbs, D. & Chen, Y. (2009) The Red List of Maples Botanic Gardens Conservation International (BGCI) The of the genus is the sycamore maple Acer pseudoplatanus, one of the most common maple species in Europe.van Gelderen, C. J. & van Gelderen, D. M. (1999). Most maples usually have easily identifiable leaves (with a few exceptions, such as Acer carpinifolium, , and ) and all share distinctive winged fruits. The closest relative of the maples is the small east Asian genus , followed by the more widespread genus (buckeyes and horse-chestnuts). is made from the sap of some maple species. It is one of the most common genera of trees in Asia. Many maple species are grown in gardens where they are valued for their autumn colour and often decorative foliage, some also for their attractive flowers, fruit, or bark.[2] Crowley (2020) Acer L. from the website Trees and Shrubs Online


Evolutionary history
The closest relative of Acer is , which only has two living species in China, but has a fossil record extending back to the middle in North America. The oldest known fossils of Acer are from the late Paleocene of and northern North America, around 60 million years old. The oldest fossils of Acer in Europe are from , dating to the late ( ~38–34 million years ago).


Morphology
Most maples or acers are trees growing to a height of . Others are shrubs less than 10 meters tall with a number of small trunks originating at about ground level. Most species are , and many are renowned for their autumn leaf colours, but a few in southern Asia and the Mediterranean region are mostly . Most are when young and are often riparian, understory, or pioneer species rather than climax overstory trees. There are a few exceptions such as , which may form monodominant forests. Many of the systems are typically dense and fibrous, inhibiting the growth of other vegetation underneath them. A few species, notably Acer cappadocicum, frequently produce , which can develop into .

Maples are distinguished by opposite arrangement. The leaves in most species are veined and lobed, with 3 to 9 (rarely to 13) veins each leading to a lobe, one of which is central or apical. A small number of species differ in having palmate compound, compound, pinnate veined or unlobed leaves. Several species, including (paperbark maple), Acer mandshuricum (Manchurian maple), Acer maximowiczianum (Nikko maple) and (three-flowered maple), have trifoliate leaves. One species, (box-elder or Manitoba maple), has pinnately compound leaves that may be simply trifoliate or may have five, seven, or rarely nine leaflets. A few, such as (Nepal maple) and Acer carpinifolium (hornbeam maple), have pinnately veined simple leaves.

Maple species, such as , may be , or polygamodioecious. The are regular, pentamerous, and borne in , , or . They have four or five , four or five about 1–6 mm long (absent in some species), four to ten about 6–10 mm long, and two or a pistil with two styles. The ovary is superior and has two , whose wings elongate the flowers, making it easy to tell which flowers are female. Maples flower in late or early spring, in most species with or just after the appearance of the leaves, but in some before the trees leaf out.Huxley, A., ed. (1992). New RHS Dictionary of Gardening. Macmillan .

Maple flowers are green, yellow, orange or red. Though individually small, the effect of an entire tree in flower can be striking in several species. Some maples are an early spring source of and for .

The distinctive are called samaras, "maple keys", "helicopters", "whirlybirds" or "polynoses". These occur in distinctive pairs each containing one seed enclosed in a "nutlet" attached to a flattened wing of fibrous, papery tissue. They are shaped to spin as they fall and to carry the seeds a considerable distance on the wind. People often call them "helicopters" due to the way that they spin as they fall. During World War II, the US Army developed a special airdrop supply carrier that could carry up to of supplies and was based on the maple seed. "Sky Hook Spirals from Plane" Popular Mechanics, December 1944, p. 75. Seed maturation is usually in a few weeks to six months after flowering, with seed dispersal shortly after maturity. However, one tree can release hundreds of thousands of seeds at a time. Depending on the species, the seeds can be small and green to orange and big with thicker seed pods. The green seeds are released in pairs, sometimes with the stems still connected. The yellow seeds are released individually and almost always without the stems. Most species require stratification in order to , and some seeds can remain dormant in the soil for several years before germinating.

The genus Acer, together with genus Dipteronia, were formerly often classified in a family of their own, the , but recent botanical consensus, including the Angiosperm Phylogeny Group system, includes them in the family ; their exclusion from Sapindaceae would leave that family . Within Sapindaceae, Acer is placed in the Hippocastanoideae. The genus is subdivided by its morphology into a multitude of sections and subsections. Molecular studies incorporating DNA sequence data from both chloroplast and nuclear genomes, aiming to resolve the internal relationships and reconstruct the evolutionairy history of the group, suggest a Late Paleocene origin for the group, appearing first in the northeastern Palearctic. Rapid lineage divergence was followed by several independent dispersals to the Nearctic and Western Palearctic regions. Fifty-four species of maples meet the International Union for Conservation of Nature criteria for being under threat of extinction in their native habitat.


Pests and diseases
The leaves are used as a food plant for the of a number of the order (see List of Lepidoptera that feed on maples). In high concentrations, caterpillars, like the greenstriped mapleworm ( Dryocampa rubicunda), can feed on the leaves so much that they cause temporary defoliation of host maple trees. are also very common sap-feeders on maples. In horticultural applications a spray will solve this.

Infestations of the Asian long-horned beetle ( Anoplophora glabripennis) have resulted in the destruction of thousands of maples and other tree species in Illinois, Massachusetts, New Jersey, New York, and Ohio in the United States and Ontario, Canada. September 18, 2003 Asian Longhorned Beetle discovered in York Region

Maples are affected by a number of diseases. Several are susceptible to Verticillium wilt caused by species, which can cause significant local mortality. Sooty bark disease, caused by species, can kill trees that are under stress due to . Death of maples can rarely be caused by and root decay. Maple leaves in late summer and autumn are commonly disfigured by "tar spot" caused by Rhytisma species and caused by species, though these diseases do not usually have an adverse effect on the trees' long-term health.Phillips, D. H. & Burdekin, D. A. (1992). Diseases of Forest and Ornamental Trees. Macmillan. .


Cultural significance
A appears on the coat of arms of Canada, and is on the . The maple is a common symbol of strength and endurance and has been chosen as the national tree of Canada. Maple leaves are traditionally an important part of military regalia, for example, the military rank insignia for generals use maple leaf symbols. There are 10 species naturally growing in the country, with at least one in each province. Although the idea of the tree as a national symbol originally hailed from the province of where the is significant, today's arboreal emblem of Canada rather refers to a generic maple. The design on the flag is an eleven-point stylization modeled after a sugar maple leaf (which normally bears 23 points).
(2025). 9781134687633, Routledge. .

It is also in the name of the Canadian ice hockey team, the Toronto Maple Leafs.

The first attested use of the word was in 1260 as "mapole", and it also appears a century later in 's , spelled as "mapul". The maple is also a symbol of , ubiquitous in the local .

A maple leaf, along with samaras, appears in the coat of arms of , a former municipality of , .

(1982). 9789517730853, Suomen Kunnallisliitto.


Uses

Horticulture
Some species of maple are extensively planted as by homeowners, businesses, and municipalities due to their fall colour, relatively fast growth, ease of transplanting, and lack of hard seeds that would pose a problem for mowing lawns. Particularly popular are (although it is considered invasive in North America), , , and . The is also occasionally used as an ornamental tree. Other maples, especially smaller or more unusual species, are popular as specimen trees.


Cultivars
Numerous maple that have been selected for particular characteristics can be propagated only by asexual reproduction such as cuttings, , budding or . (Japanese maple) alone has over 1,000 cultivars, most selected in Japan, and many of them no longer propagated or not in cultivation in the . Some delicate cultivars are usually grown in pots and rarely reach heights of more than 50–100 cm.


Bonsai
Maples are a popular choice for the art of . ( Acer palmatum), ( A. buergerianum), ( A. ginnala), ( A. campestre) and Montpellier maple ( A. monspessulanum) are popular choices and respond well to techniques that encourage leaf reduction and ramification, but most species can be used.


Collections
Maple collections, sometimes called aceretums, occupy space in many gardens and around the world including the "five great W's" in : Wakehurst Place Garden, Westonbirt Arboretum, Windsor Great Park, Winkworth Arboretum and . In the , the aceretum at the Harvard-owned in Boston is especially notable. In the number of species and cultivars, the in , is the largest in the world.


Commercial uses
Maples are important as sources of syrup and wood. Dried wood is often used for the smoking of food. Charcoal from maples is an integral part of the Lincoln County Process used to make Tennessee whiskey. They are also cultivated as and have benefits for and .


Timber
Some of the larger maple species have valuable , particularly Sugar maple in North America and Sycamore maple in Europe. Sugar maple wood—often known as "hard maple"—is the wood of choice for pins, bowling alley lanes, pool and , and . Maple wood is also used for the manufacture of wooden , though less often than ash or due to the tendency of maple bats to shatter if they do break. The maple bat was introduced to Major League Baseball (MLB) in 1998 by founder Sam Holman. Today it is the standard maple bat most in use by professional baseball. Sam Holman Maple Baseball Bats Maple is also commonly used in archery as the core material in the limbs of a due to its stiffness and strength.

Maple wood is often graded based on physical and aesthetic characteristics. The most common terminology includes the grading scale from common #2; which is unselected and often used for craft woods; common #1, used for commercial and residential buildings; clear; and select grade, which is sought for fine woodworking.

Some maple wood has a highly decorative , variously known as , , and . This condition occurs randomly in individual trees of several species and often cannot be detected until the wood has been sawn, though it is sometimes visible in the standing tree as a rippled pattern in the bark.

These select decorative wood pieces also have subcategories that further filter the aesthetic looks. Crotch wood, bees wing, cats paw, old growth and mottled are some terms used to describe the look of these decorative woods. Wood Terms and Examples

Maples have a long history of use for furniture production in the United States.

(1965). 9780517037355, Random House, Inc.. .
The people would produce a purple dye from maple bark, which they used to dye cloth.
(2025). 9780806109237, University of Oklahoma Press. .


Tonewood
Maple is considered a , or a wood that carries sound waves well, and is used in numerous musical instruments. Maple is harder and has a brighter sound than , which is another major tonewood used in instrument manufacturing.

The back, sides, and neck of most , , , and are made from maple.

necks are commonly made from maple, having good dimensional stability. The necks of the Fender and were originally an entirely maple one piece neck, but later were also available with fingerboards. desired an all maple guitar, but due to the weight of maple, only the tops of Gibson's Les Paul guitars are made from carved maple, often using quilted or flamed maple tops. Due to its weight, very few guitars are made entirely from maple, but many guitars have maple necks, tops or veneers.

Maple is also often used to make and sometimes for other woodwind instruments like maple recorders.

Many are made from maple. From the 1970s to the 1990s, maple drum kits were a vast majority of all drum kits made, but in recent years, has become popular for drums once again. Some of the best drum-building companies use maple extensively throughout their mid-pro range. Maple drums are favored for their bright resonant sound.

(1997). 9780879304768, Hal Leonard Corporation. .
Certain types of drum sticks are also made from maple.


Agriculture
During late winter to early spring in northeastern , when the night-to-day temperatures change from freezing to thawing, maple trees may be tapped for sap to manufacture . The sap is sent via tubing to a where it is boiled to produce syrup or made into or . It takes about of sugar maple sap to make of syrup. While any Acer species may be tapped for syrup, many do not have sufficient quantities of sugar to be commercially useful, whereas ( A. saccharum) are most commonly used to produce maple syrup. Québec, Canada is a major producer of maple syrup, an industry worth about 500 million annually.

Also, as these trees are a major source of in early spring before many other plants have flowered, maple flowers are a source of foraging for that play a commercially important role in general agriculture and in natural habitats.


Pulpwood
Maple is used as . The fibers have relatively thick walls that prevent collapsing upon drying. This gives good bulk and opacity in . Maple also gives paper good printing properties.


Tourism
Many maples have bright , and many countries have leaf-watching traditions. The sugar maple ( ), whose leaves turn brilliant orange, is the primary contributor to fall "" in north-eastern . In , the custom of viewing the changing colour of maples in the autumn is called . Nikkō and are particularly favoured destinations for this activity. In Korea, the same viewing activity is called danpung-nori and the and mountains are among the best-known destinations.


Gallery
File:台灣三角楓 Acer buergerianum var. formosanum 20220411100147 08.jpg| Acer buergerianum var. formosanum leaves and fruit Image:Acer cappadocicum spring.jpg| Acer cappadocicum (Cappadocian maple) Image:Acer carpinifolium.jpg| Acer carpinifolium leaves Image:Acer ginnala.jpg| foliage Image:Bi-colored Maple Tree.jpg| Acer grandidentatum (bigtooth maple) in autumn colour Image:Paperbark Maple Acer griseum Leaves Closeup 2856px.jpg| (paperbark maple) Image:Acer laevigatum 3.jpg| leaves and fruit Image:Acer macrophyllum 0304.jpg| Acer macrophyllum flowers and young leaves File:青楓 Acer serrulatum 20210419095802 02.jpg| The Chinese name "Qingfeng" comes from the bark color of the new branches and young trunk which are green. Image:TenryujiMomiji.jpg| trees and bamboo in Japan Image:Red maple leaf.jpg| leaf in Image:Maple leaf Fcb981.JPG| leaf Image:Helicopter leaves.jpg| (Norway maple) samaras File:Acer rubrum 1-eheep (5097479399).jpg| leaves File:Red maple.png| tree in autumn Image:Acer sempervirens leaves.jpg| Acer sempervirens foliage Image:Autumn Blaze Maple Foliage.jpg| Acer × freemanii 'Autumn Blaze' (a cross between and


See also
  • List of Acer species
  • List of Award of Garden Merit maples
  • List of Danish Acers
  • Mazer – a drinking vessel made from maple wood
  • List of foods made from maple


Citations
(1999). 9780585254579, Timber Press.


General bibliography

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