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Rye ( Secale cereale) is a grown extensively as a , a and a crop. It is grown principally in an area from Eastern and Northern Europe into Russia. It is much more tolerant of cold weather and poor soil than other cereals, making it useful in those regions; its vigorous growth suppresses weeds and provides abundant forage for animals early in the year. It is a member of the wheat tribe () which includes the cereals and . It is likely that rye arrived in Europe as a , meaning that it was a minor admixture in wheat as a result of Vavilovian mimicry, and was only later cultivated in its own right.

Rye grain is used for , , , and animal . In Scandinavia, rye was a staple food in the Middle Ages, and rye remains a popular food in the region. Europe produces around half of the world's rye; relatively little is traded between countries. A wheat-rye hybrid, , combines the qualities of the two parent crops and is produced in large quantities worldwide. In European folklore, the Roggenwolf ("rye wolf") is a carnivorous corn demon or Feldgeister]].


Origins
The rye genus is in the tribe , which contains other such as ( Hordeum) and ( Triticum).

The generic name Secale, related to Italian segale and French seigle meaning "rye", is of unknown origin but may derive from a Balkan language. The English name rye derives from Old English ryge, related to Dutch rogge, German Roggen, and Russian рожь , again all with the same meaning.

Rye is one of several cereals that grow wild in the , central and eastern and adjacent areas. Evidence uncovered at the site of Tell Abu Hureyra in the valley of northern suggests that rye was among the first cereal crops to be systematically cultivated, around 13,000 years ago. However, that claim remains controversial; critics point to inconsistencies in the dates, and identifications based solely on grain, rather than on .

Domesticated rye occurs in small quantities at a number of sites in Asia Minor (, now Turkey), such as the Pre-Pottery Neolithic B Can Hasan III near Çatalhöyük, but is otherwise absent from the archaeological record until the of central Europe, c. 1800–1500 BCE.

(2025). 9780199549061, Oxford University Press. .

It is likely that rye was brought westwards from Asia Minor as a , meaning that it was a minor admixture in wheat as a result of Vavilovian mimicry, and was only later cultivated in its own right. Archeological evidence of this grain has been found in contexts along the and the and in Ireland and Britain. page 26. The Roman naturalist Pliny the Elder was dismissive of a grain that may have been rye, writing that it "is a very poor food and only serves to avert starvation".

(1981). 9780521237932, Cambridge University Press. .
He said it was mixed with "to mitigate its bitter taste, and even then is most unpleasant to the stomach".


Description
Rye is a tall grass grown for its seeds; it can be an or a . Depending on environmental conditions and variety it reaches in height. Its leaves are blue-green, long, and pointed. The seeds are carried in a curved head or spike some long. The head is composed of many , each of which holds two small flowers; the spikelets alternate left and right up the head.

File:Illustration Secale cereale0.jpg|Botanical illustration File:Secale cereale - cereal rye 2 - Steve Hurst USDA-NRCS PLANTS Database.jpg|Seed in File:Secale cereale - cereal rye - Steve Hurst USDA-NRCS PLANTS Database.jpg|Different types of grains File:Rye grains rotated (cropped).jpg|The seeds of rye are some 7 or 8 mm long, much larger and less round than .


Cultivation
Since the , people have cultivated rye widely in and . It serves as the main cereal in most areas east of the France–Germany border and north of . In Southern Europe, it was cultivated on marginal lands.

Rye grows well in much poorer soils than those necessary for most cereal grains. Thus, it is an especially valuable crop in regions where the soil has or . Rye plants withstand cold better than other small grains, surviving snow cover that would kill winter wheat. Winter rye is the most popular: it is planted and begins to grow in autumn. In spring, the plants develop rapidly. This allows it to provide spring grazing, at a time when spring-planted wheat has only just germinated.

The physical properties of rye affect attributes of the final food product such as seed size, surface area, and porosity. The surface area of the seed directly correlates to the drying and heat transfer time. Smaller seeds have increased heat transfer, which leads to lower drying time. Seeds with lower porosity lose water more slowly during the process of drying.

Rye is like wheat with a combine harvester, which cuts the plants, and the grain, and releases the to the field where it is later pressed into bales or left as soil amendment. The resultant grain is stored in local or transported to regional and combined with other lots for storage and distant shipment. Before the era of mechanised agriculture, rye harvesting was a task performed with or .

(1988). 9780300042658, Yale University Press (YUP). .
(2025). 9780191025150, Oxford University Press (OUP). .


Agroecology
Winter rye is any breed of rye planted in the autumn to provide ground cover for the winter. It grows during warmer days of the winter when sunlight temporarily warms the plant above freezing, even while there is general snow cover. It can be used as a cover crop to prevent the growth of weeds.
(2025). 9781560229575, Haworth Press.

Rye grows better than any other cereal in heavy and light sandy soil, and infertile or drought-affected soils. It can tolerate pH between 4.5 and 8.0, but soils having pH 5.0 to 7.0 are best suited for rye cultivation. Rye grows best in fertile, well-drained or clay-loam soils.

(2025). 9781848263680, Publishers. .
As for temperature, the crop can thrive in subzero environments, assisted by the production of antifreeze polypeptides (different from those produced by some fish and insects) by the leaves of winter rye.

Rye is a common, of fields. If allowed to grow and mature, it may cause substantially reduced prices (docking) for harvested wheat.


Pests and diseases
Pests including the nematode Ditylenchus dipsaci and a variety of herbivorous insects can seriously affect plant health.
(1991). 9780442308308, Van Nostrand Reinhold/AVI. .

Rye is highly susceptible to the fungus. ergot , online medical dictionary ergot, Dorland's Medical Dictionary Consumption of ergot-infected rye by humans and animals results in , which causes convulsions, miscarriage, necrosis of digits, hallucinations and death. Historically, damp northern countries that depended on rye as a staple crop were subject to periodic epidemics. Modern grain-cleaning and milling methods have practically eliminated ergotism, but it remains a risk if food safety vigilance breaks down.

After an absence of 60 years, stem rust ( Puccinia graminis f. sp. tritici) has returned to in the 2020s. Areas affected include , (), , and .


Production
+ Rye production
3,124,200
2,533,220
1,700,000
780,000
601,130
518,640
12,680,515

In 2023, world production of rye was 12.7 million , led by Germany with 25% of the total, and Poland and Russia as major secondary producers (table).


Nutrition
Raw rye is 11% water, 76% , 10% protein, and 2% (table). A reference amount of provides of , and is a rich source (20% or more of the , DV) of , , such as and niacin (each at 25% DV), and several dietary minerals, including manganese (130% DV), zinc, phosphorus, and magnesium (26-27% DV) (table).


Health effects
According to and the U.S. Food and Drug Administration, consuming at least per day of rye or per serving of can lower levels of blood cholesterol, a risk factor for cardiovascular diseases.

Eating whole-grain rye, as well as other high-fiber grains, improves regulation of (i.e., reduces blood glucose response to a meal). Consuming containing rye over weeks to months also improved cholesterol levels and glucose regulation.


Health concerns
Like wheat, barley, and their hybrids and derivatives, rye contains and related prolamines, which makes it an unsuitable grain for consumption by people with gluten-related disorders, such as , non-celiac gluten sensitivity, and , among others. Nevertheless, some wheat allergy patients can tolerate rye or barley.


Uses

Food and drink
Rye grain is refined into a high in but low in and rich in soluble fiber. are phenolic lipids present in high amounts in the bran layer (e.g. , testa and layers) of wheat and rye (0.1–0.3% of dry weight). , including , is made using rye flour and is a widely eaten food in Northern and Eastern Europe. In Scandinavia, rye is widely used to make (Knäckebröd); in the Middle Ages it was a in the region, and it remains popular in the 21st century.

Rye grain is used to make alcoholic drinks, such as and . The traditional cloudy and sweet-sour low-alcohol beverage is fermented from rye bread or rye flour and malt.

(2025). 9781849731614, Royal Society of Chemistry.

Oval rye pumpkin bread 2024-03-10 08.jpg| Naturaplan Bio Pumpernickel.png| IKEA knäckebröd.JPG|Swedish rye (Knäckebröd) File:Sultsina.png|, a traditional dish made of unleavened rye dough and a farina filling Old Overholt Rye Whiskey bottle and tumbler.jpg|Rye whiskey Terrapin Brewing Co. Rye Squared Imperial Pale Ale.jpg|Rye beer


Other uses
Rye is a useful crop in cool climates; it grows vigorously and provides plentiful for grazing animals, or to improve the soil. It forms a good in winter with its rapid growth and deep roots.

Rye straw is used as livestock bedding, despite the risk of ergot poisoning. It is used on a small scale to make such as . More recently it has found uses as a raw material for bioconversion to products such as the sweetener .

(2025). 9780128114131, Elsevier.

Rye flour is boiled with red iron oxide pigments and some additives to make traditional paint, widely used as a house paint in Sweden and other Scandinavian countries.


Production of hybrids
Plant breeders, starting in the 19th century in Germany and Scotland, but mainly from the 1950s, worked to develop a hybrid cereal with the best qualities of wheat and rye, now called . Modern triticales are with six sets of chromosomes; they are used to produce millions of tons of cereal annually.

Varieties of rye hold much genetic diversity,

(2025). 9783030833831, .
which cites which can be used to improve other crops such as wheat. For example, the pollination abilities of wheat can be improved by the addition of the rye chromosome 4R; this increases the size of the wheat and the amount of pollen. The chromosome is the source of many crop disease resistance genes. Varieties such as Petkus, Insave, Amigo, and Imperial have donated 1R-originating resistance to wheat. AC Hazlet rye is a medium-sized winter rye with resistance to both lodging and shattering. Rye was the of Sr31 – a stem rust resistance gene – introgressed into wheat.

The characteristics of S. cereale have been combined with another perennial rye, , to produce , which has the beneficial characteristics of each. The hybrid rye can be grown in harsh environments and on poor soil. It provides improved forage with digestible fiber and protein.


In human culture
In European folklore, the Roggenwolf ("rye wolf") is a carnivorous corn demon or , a field spirit shaped like a .
(2025). 9783937715384, Marix-Verlag.
The Roggenwolf steals children and feeds on them.
(2025). 142124778X, Elibron Classics. 142124778X
The last grain heads are often left at their place as a sacrifice for the agricultural spirits.
(2025). 9783937715391, Marix-Verlag.

In contrast, the Roggenmuhme or Roggenmutter ("rye aunt" or "rye mother") is an anthropomorphic female corn demon with fiery fingers. Her bosoms are filled with tar and may end in tips of iron. Her bosoms are also long, and as such must be thrown over her shoulders when she runs. The Roggenmuhme is completely black or white, and in her hand she has a birch or whip from which lightning sparks. She can change herself into different animals, such as snakes, turtles, and frogs.

(2025). 9783955627980, Bremen University Press.

The classical scholar Carl A. P. Ruck writes that the Roggenmutter was believed to go through the fields, rustling like the wind, with a pack of rye wolves running after her. They spread ergot through the sheaves of harvested rye. According to Ruck, they then lured children into the fields to nurse on the infected grains "like the iron teats of the Roggenmutter". The enlarged reddish ergot-infected grains were known as Wulfzähne (wolf teeth).


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