Barley (Hordeum vulgare), a member of the Poaceae, is a major cereal grain grown in temperate climates globally. It was one of the founder crops; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering and making it much easier to harvest. Its use then spread throughout Eurasia by 2000 BC. Barley prefers relatively low temperatures and well-drained soil to grow. It is relatively tolerant of drought and soil salinity but is less winter-hardy than wheat or rye.
In 2022, barley was fourth among grains in quantity produced, 155 million , behind maize, wheat, and rice. Globally, 70% of barley production is used as animal feed, while 30% is used as a source of fermentable material for beer, or further Distillation into whisky, and as a component of various foods. It is used in soups and stews and in barley bread of various cultures. Barley grains are commonly made into malt using a traditional and ancient method of preparation. In English folklore, John Barleycorn personifies the grain and the alcoholic beverages made from it. English pub names such as The Barley Mow allude to its role in the production of beer.
The genome of barley was sequenced in 2012 by the International Barley Genome Sequencing Consortium and the UK Barley Sequencing Consortium. The genome is organised into seven pairs of nuclear DNA (recommended designations: 1H, 2H, 3H, 4H, 5H, 6H and 7H), and one mitochondrial and one chloroplast DNA, with a total of 5000 Mbp. Details of the genome are freely available in several barley databases.
Domestication changed the Plant morphology of the barley grain substantially, from an elongated shape to a more rounded spherical one. Wild barley has distinctive , , and regulators with potential for resistance to abiotic or ; these may help cultivated barley to adapt to climatic changes. Wild barley has a brittle spike; upon maturity, the separate, facilitating seed dispersal. Domesticated barley has nonshattering spikelets, making it much easier to harvest the mature ears. The nonshattering condition is caused by a mutation in one of two Genetic linkage genes known as Bt1 and Bt2; many possess both mutations. The nonshattering condition is recessive, so varieties of barley that exhibit this condition are homozygous for the mutant allele. Domestication in barley is followed by the change of key phenotype at the genetic level.
The wild barley found currently in the Fertile Crescent may not be the progenitor of the barley cultivated in Eritrea and Ethiopia, indicating that it may have been domesticated separately in eastern Africa.
Rations of barley for workers appear in Linear B tablets in Mycenaean contexts at Knossos and at Mycenaean Pylos. In mainland Greece, the ritual significance of barley possibly dates back to the earliest stages of the Eleusinian Mysteries. The preparatory kykeon or mixed drink of the initiates, prepared from barley and herbs, mentioned in the Homeric hymn to Demeter. The goddess's name may have meant "barley-mother", incorporating the ancient Cretan word δηαί (dēai), "barley". The practice was to dry the barley groats and roast them before preparing the porridge, according to Pliny the Elder's Natural History.Pliny the Elder. Natural History, xviii.72. Tibetan barley has been a staple food in Tibetan cuisine since the fifth century AD. This grain, along with a cool climate that permitted storage, produced a civilization that was able to raise great armies. It is made into a flour product called tsampa that is still a staple in Tibet. In medieval Europe, bread made from barley and rye was peasant food, while wheat products were consumed by the upper classes.
In traditional taxonomy, different forms of barley were classified as different species based on morphological differences. Two-row barley with shattering spikes (wild barley) was named Hordeum spontaneum. Two-row barley with nonshattering spikes was named as H. distichon, six-row barley with nonshattering spikes as H. vulgare (or H. hexastichum), and six-row with shattering spikes as H. agriocrithon. Because these differences were driven by single-gene mutations, coupled with cell biology and molecular evidence, most recent classifications treat these forms as a single species, H. vulgare.
Like other cereals, barley is typically planted on Tillage land. Seed was sowing, but in developed countries is usually seed drill. As it grows it requires soil nutrients (nitrogen, phosphorus, potassium), often supplied as fertilizers. It needs to be monitored for pests and diseases, and if necessary treated before these become serious. The stems and ears turn yellow when ripe, and the ears begin to droop. Traditional harvesting was by hand with or ; in developed countries, harvesting is mechanised with combine harvesters.
Serious fungal diseases of barley include powdery mildew caused by Blumeria graminis, leaf scald caused by Rhynchosporium secalis, barley rust caused by Puccinia hordei, crown rust caused by Puccinia coronata, various diseases caused by Cochliobolus sativus, Fusarium],
and stem rust ( Puccinia graminis).
Bacterial diseases of barley include bacterial blight caused by Xanthomonas campestris pv. translucens.
Barley is susceptible to several viral diseases, such as barley mild mosaic bymovirus. Some viruses, such as barley yellow dwarf virus, Disease vector by the rice root aphid, can cause serious crop injury.
For durable disease resistance, quantitative resistance is more important than qualitative resistance. The most important have corresponding resistance gene regions on all chromosomes of barley.
A large number of are available for breeding of resistance to leaf rust, powdery mildew, Rhynchosporium secalis, Pyrenophora teres f. teres, Barley yellow dwarf virus, and the Barley yellow mosaic virus complex.
With a long history of cultivation in the Middle East, barley is used in a wide range of traditional Arab cuisine, Assyrian cuisine, Israelite, Kurdish cuisine, and Persian cuisine foodstuffs including keşkek, kashk, and murri. Barley soup is traditionally eaten during Ramadan in Saudi Arabia. Cholent (in Hebrew) is a traditional Jewish stew often eaten on the Shabbat, in numerous recipes by both Mizrahi Jews and Ashkenazi Jews; its original form was a barley porridge.
In Eastern Europe and Central Europe Europe, barley is used in soups and stews such as ričet. In Africa, where it is a traditional food plant, it has the potential to improve nutrition, boost food security, foster rural development, and support sustainable landcare.
The six-row variety bere is cultivated in Orkney, Shetland, Caithness and the Western Isles of the Scottish Highlands and Islands. When milled into beremeal, it is used locally in bread, , and the traditional beremeal bannock.
In Japanese cuisine cuisine, barley is mixed with rice and steamed as mugimeshi. The naval surgeon Takaki Kanehiro introduced it into institutional cooking to combat beriberi, endemic in the armed forces in the 19th century. It became standard prison fare, and remains a staple in the Japan Self-Defense Forces.
Nonalcoholic drinks such as barley water and roasted barley tea have been made by boiling barley in water. In Italy, roasted barley is sometimes used as coffee substitute, caffè d'orzo (barley coffee).
In English folklore, the figure of John Barleycorn in the folksong of the same name is a personification of barley, and of the alcoholic beverages made from it: beer and whisky. In the song, John Barleycorn is represented as suffering attacks, death, and indignities that correspond to the various stages of barley cultivation, such as reaping and malting; but he is revenged by getting the men drunk: "And little Sir John and the nut-brown bowl / Proved the strongest man at last." The folksong "Elsie Marley" celebrates an alewife of County Durham with lines such as "And do you ken Elsie Marley, honey? / The wife that sells the barley, honey". The antiquary Cuthbert Sharp records that Elsie Marley was "a handsome, buxom, bustling landlady, and brought good custom to the ale house by her civility and attention."
English pub names such as The Barley Mow, John Barleycorn, Malt Shovel, and Mash Tun allude to barley's role in the production of beer.
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+ Barley in Egyptian hieroglyphs jt ideogram jt spelling šma ideogram
Taxonomy and varieties
Two-row and six-row barley
Hulless barley
23.4 14.4 11.3 11.2 10.0 8.5 7.4 7.0 154.9
Production
Cultivation
Scotland, 2009
File:Cessna188AGWagonZKCSE.jpg|Spraying barley for rust fungus,
New Zealand, 1979
File:In The Barley Harvest.jpg|Traditional barley harvest by hand with , England, c. 1886.
Photo Peter Henry Emerson
File:Claas Dominator 85 harvesting winter barley at Ostrittrum.jpg|Harvesting winter barley with a combine harvester, Germany, 2017
Pests and diseases
Food
Preparation
Nutrition
Health implications
Uses
Beer, whisky, and soft drinks
roasted barley tea
Animal feed
Other uses
Culture and folklore
Sources
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