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The Maleae (incorrectly Pyreae) are the apple tribe in the rose family, . The group includes a number of plants bearing commercially important , such as and , while others are cultivated as ornamentals. Older taxonomies separated some of this group as tribe Crataegeae,G. K. Schulze-Menz 1964. Reihe Rosales. in A. Engler's Syllabus der Pflanzenfamilien mit besonderer Berücksichtigung der Nutzpflanzen nebst einer Übersicht über die Florenreiche und Florengebiete der Erde, Gebrüder Borntraeger, Berlin as the Cydonia group (a tentative placement), or some genera were placed in family .

The tribe consists exclusively of and small . Most have , a type of that does not occur in other Rosaceae. All except (with 15 chromosomes) have a basal count of 17, instead of 7, 8, or 9 as in the other Rosaceae.

There are approximately 28 genera that contain about 1100 species worldwide, with most species occurring in the temperate Northern Hemisphere.


Current classification

Core members of the group
A traditional circumscription of Maleae includes the following genera:

intergeneric hybrids:Stace, C. A. 1975. Hybridization and the flora of the British Isles. Academic Press, London.

  • × Amelasorbus
  • × Malosorbus
  • × Sorbaronia

and :
+ Pyrocydonia ( Pirocydonia)


Tribe Crataegeae
A recent taxonomic treatment includes the following genera in Maleae that were earlier separated as tribe Crataegeae (or as intertribe hybrids):Potter, D.; Eriksson, T.; Evans, R.C.; Oh, S.H.; Smedmark, J.E.E.; Morgan, D.R.; Kerr, M.; Robertson, K.R.; Arsenault, M.P.; Dickinson, T.A.; Campbell, C.S. (2007). Phylogeny and classification of Rosaceae. Plant Systematics and Evolution. 266(1–2): 5–43.

intergeneric (including intertribal) hybrids:

  • × Crataemespilus
  • × Crataegosorbus
  • × Sorbocotoneaster

and the graft hybrid:

  • + Crataegomespilus


Former members of family Quillajaceae
The following genera were previously placed in tribe Quillajeae in Rosaceae, or in family Quillajaceae. Their fruit are dry capsules, not pomes.


The Cydonia group
The Cydonia group within the Maloid Rosaceae was a tentative grouping of pome-fruited genera with many ovules (rather than just two) per carpel. The genera involved were:

It is not yet clear whether this group is monophyletic within the Maleae. Molecular data indicate a close relationship between Cydonia and Pseudocydonia. Multiple ovules per carpel also occur in Kageneckia, a non-pome-bearing genus. DNA analysis, but not nuclear DNA, shows a tight relationship between Cydonia and Dichotomanthes, a non-pome-bearing genus.

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