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   » » Wiki: Cypripedium
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Cypripedium is a of 58 species and of hardy ; it is one of five genera that together compose the of lady's slipper orchids (). They are widespread across much of the Northern Hemisphere, including most of and (Walid, Nemer & Rebbas, Khellaf & Krouchi, Faiza. (2019). Découverte de Cypripedium calceolus (Orchidaceae) au Djurdjura (Algérie), nouvelle pour l’Afrique du Nord. Flora Mediterranea. 29. 207-214. 10.7320/FlMedit29.207.) (one species), , , , the , , and . They are most commonly known as slipper orchids, lady's slipper orchids, or ladyslippers; other common names include moccasin flower, camel's foot, squirrel foot, steeple cap, Venus' shoes, and whippoorwill shoe. An abbreviation used in trade journals is " Cyp." The genus name is derived from Κύπρις (), an early reference in Greek myth to , and πέδιλον (), meaning "sandal".

Most of Cypripedium grow in temperate and subtropical climates, but some species grow in the in and , which is an unusually cold habitat for orchids. Other species occur well into tropical areas such as and .

Some of the northern species can withstand extreme cold, growing under the snow and blooming when the snow melts. But, in the wild, some have become rare and close to extinction, due to an ever shrinking natural habitat and over-collection, people prizing the flowers for their beauty. Several species are legally protected in some regions. In the late 20th century, only a single known plant of Cypripedium calceolus survived in Britain.


Characteristics
The Cypripedium are terrestrial and, as with most terrestrial orchids, the is short and robust, growing in the uppermost soil layer. The rhizome grows annually with a growth bud at one end and dies off at the other end. The grows from the bud at the tip of the rhizome. Most slipper orchids have an elongate erect stem, with growing along its length. But the mocassin flower or pink lady's slipper ( Cypripedium acaule) has a short underground stem with leaves springing from the soil. The often hairy leaves can vary from ovate to elliptic or , folded (plicate) along their length. The stems lack .

The is . It can carry one to twelve , as in Cypripedium californicum. But most species have one to three flowers. There are three , with, in most species, the two lateral ones more or less fused. The flower has three acute with the third a striking slipper-shaped lip, which is lowermost. The sepals and the petals are usually similarly colored, with the lip in a different color. But variations on this theme occur. The aspect of the lip of different species can vary a great deal. As with all orchids, it is specially constructed to attract , which it temporarily. The flowers show a column with a unique shield-like staminode. The ovary is 3-locular (with three chambers).


Taxonomy
Comparison between a DNA-analysis and the morphological characteristics in this genus has shown that there is a high degree of divergence between the two, probably due to long periods of isolation or extinction of intermediate forms. The Eurasian species with yellow or red flowers form a distinct group from the North American species with yellow flowers. The Mexican Pelican Orchid ( Cypripedium irapeanum) and the California lady's slipper ( Cypripedium californicum) are probably the first diverging line. They share several similarities with their sister group .


Species and natural hybrids
There are 58 currently recognized species and (naturally occurring hybrids) recognized in this genus, as of May 2014:


Subgenus Cypripedium
Cypripedium plectrochilumN Myanmar to SC China
Cypripedium yatabeanumRussian Far East to N & NC Japan, Aleutian Islands to SW Alaska
Cypripedium candidum – Small white lady slipperSE Canada, NC & E USA
Cypripedium cordigerum – Heart-Lip lady's slipperN Pakistan to Himalaya, S Tibet
Cypripedium farreriChina
Cypripedium fasciolatumChina
Cypripedium henryi – Henry's lady's slipperC China
Cypripedium kentuckiense – Kentucky lady's slipper, Southern lady's slipperC & E USA
Cypripedium montanum – Large lady's slipper, Mountain lady's slipper, White lady's slipper, Moccasin flowerAlaska to California
Cypripedium parviflorum Salisb. – (Greater) yellow lady's slipper, moccasin flower, or hairy yellow ladyslipperCanada, E USA
Cypripedium segawaiEC Taiwan
Cypripedium shanxienseChina to N Japan
Cypripedium formosanum – Formosa lady's slipperC Taiwan
Cypripedium franchetii – Franchet's lady's slipperC & SC China
Cypripedium himalaicumSE Tibet to Himalaya
Cypripedium ludlowiiSE Tibet
Cypripedium macranthos – Large-flowered lady's slipperE Belarus to temperate E Asia
Cypripedium taibaienseChina
Cypripedium taiwanalpinumTaiwan
Cypripedium tibeticumSikkim to C China
Cypripedium yunnanenseSE Tibet, China
Cypripedium debile – Frail lady's slipperJapan, Taiwan, China
Cypripedium palangshanenseChina
Cypripedium forrestiiChina
Cypripedium micranthumChina
Cypripedium fargesiiChina
Cypripedium lentiginosumChina
Cypripedium lichiangense S.C.Chen & P.J.CribbChina (SW Sichuan, NW Yunnan), NE Myanmar
Cypripedium malipoense S.C.Chen & Z.J.LiuYunnan, China South-Central
Cypripedium margaritaceum – Pearl-white lady's slipperChina
Cypripedium sichuanenseChina
Cypripedium wumengenseChina (NE. Yunnan)


Subgenus Irapeana
Cypripedium conzattianumMexico (Colima)
Cypripedium gomezianumMexico (Colima)
Cypripedium luzmarianumMexico (Jalisco, Michoacán)
Cypripedium molleMexico
Cypripedium dickinsonianumMexico (S Chiapas) to Guatemala
Cypripedium irapeanum – Pelican Orchid, Irapeao lady's slipperMexico to Honduras
Cypripedium susanaeMexico (Nayarit)
Cypripedium passerinum – Franklyn's lady's slipper, small white Northern lady's slipper, sparrow's egg lady's slipperAlaska to Canada, Montana
Cypripedium reginae Walter – Large white lady's slipper, Queen's lady's slipper, showy lady's slipperC & E Canada, E. USA
Cypripedium wardiiSE. Tibet, China


Natural Hybrids
Alaska
E Canada, NC & NE USA
Russian Far East
W Canada, NW USA
Manitoba, Canada
Russia to Korea
China


Uses
The genus has a long history of use, dating back 2,500 years to the Far East, where they were used medicinally.


Conservation
Several orchid species thought to be extinct in the United Kingdom including one native species in this genus have been found in habitat and are currently the subject of aggressive conservation efforts to protect and restore these showy plants to their native ranges.


Awards
The following have received the Royal Horticultural Society's Award of Garden Merit:-
  • Cypripedium formosanum
  • Cypripedium Hank Small gx
  • Cypripedium Michael gx
  • Cypripedium reginae
  • Cypripedium Sabine gx

  • Phillip Cribb & Peter Green (1997). The Genus Cypripedium (a botanical monograph). Kew Royal Botanic Gardens, Timber Press
  • Pridgeon, A.M.; Cribb, P.J.; Chase, M.W. & F. N. Rasmussen (1999): Genera Orchidacearum Vol.1, page: 114 ff., Oxford U. Press.


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