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In a multicellular , an organ is a collection of tissues joined in a structural unit to serve a common function.

(2026). 9780071283663, McGraw-Hill Higher Education.
In the hierarchy of life, an organ lies between tissue and an . Tissues are formed from same type cells to act together in a function. Tissues of different types combine to form an organ which has a specific function. The intestinal wall for example is formed by epithelial tissue and smooth muscle tissue.
(2026). 9780199141951, Oxford University Press.
Two or more organs working together in the execution of a specific body function form an organ system, also called a biological system or body system.

An organ's tissues are broadly classified into , the functional tissue, and stroma, the structural tissue with supportive, connective, or functions. For example, the tissue that produces is the , while the stroma includes the that innervate the parenchyma, the that oxygenate and nourish it and remove wastes, and the connective tissues that provide structure, placement, and anchoring. The primary tissues that form an organ generally have common origins, often arising from the same . Organs are present in most multicellular . In single-celled organisms such as , the functional analogue of an organ is an . In plants, there are three main organs. The number of organs in any organism depends on the definition used. There are approximately 79 organs in the human body; the exact number remains debated.


Animals
Except for , including humans have a variety of . These specific systems are widely studied in . The functions of these organ systems often share significant overlap. For instance, the and both operate via a shared organ, the . For this reason, the two systems are combined and studied as the neuroendocrine system. The same is true for the musculoskeletal system because of the relationship between the and .


Viscera
In the study of , viscera (: viscus) refers to the internal organs of the , , and . The abdominal organs may be classified as solid organs or hollow organs. The solid organs are the , , , , and . The hollow organs of the abdomen are the , , , , and . In the , the is a hollow, muscular organ. is the study of the viscera. The term "visceral" is contrasted with the term "", meaning "of or relating to the wall of a body part, ". The two terms are often used in describing a membrane or piece of connective tissue, referring to the opposing sides.


Origin and evolution
The organ level of organisation in can be first detected in and the more derived , i.e. the . The less-advanced (i.e. , , and ) do not show unification of their tissues into organs.

More complex animals are composed of different organs, which have evolved over time. For example, the liver and heart evolved in the about 550–500 million years ago, while the gut and brain are even more ancient, arising in the ancestor of vertebrates, insects, , and worms about 700–650 million years ago.

Given the ancient origin of most vertebrate organs, researchers have looked for model systems, where organs have evolved more recently, and ideally have evolved multiple times independently. An outstanding model for this kind of research is the , which has evolved more than 100 times independently in vertebrates, has evolved relatively recently in some lineages, and exists in intermediate forms in extant taxa. Studies on the evolution of the placenta have identified a variety of genetic and physiological processes that contribute to the origin and evolution of organs, these include the re-purposing of existing animal tissues, the acquisition of new functional properties by these tissues, and novel interactions of distinct tissue types.


Plants
The study of plant organs is covered in . Organs of can be divided into vegetative and reproductive. Vegetative plant organs include , , and . The reproductive organs are variable. In , they are represented by the , and . In , the organ that bears the reproductive structures is called a . In other divisions () of plants, the reproductive organs are called , in , or simply gametophores in . Common organ system designations in plants include the differentiation of shoot and root. All parts of the plant above ground (in non-), including the functionally distinct leaf and flower organs, may be classified together as the shoot organ system.

The vegetative organs are essential for maintaining the life of a plant. While there can be 11 organ systems in animals, there are far fewer in plants, where some perform the vital functions, such as , while the reproductive organs are essential in . However, if there is asexual vegetative reproduction, the vegetative organs are those that create the new generation of plants (see ).


Society and culture
Many societies have a system for , in which a living or deceased donor's organ are into a person with a failing organ. The transplantation of larger solid organs often requires immunosuppression to prevent organ rejection or graft-versus-host disease.

There is considerable interest throughout the world in creating laboratory-grown or .


Organ transplants
Beginning in the 20th century, organ transplants began to take place as scientists knew more about the anatomy of organs. These came later in time as procedures were often dangerous and difficult. Both the source and method of obtaining the organ to transplant are major ethical issues to consider, and because organs as resources for transplant are always more limited than demand for them, various notions of justice, including distributive justice, are developed in the ethical analysis. This situation continues as long as transplantation relies upon organ donors rather than technological innovation, testing, and industrial manufacturing.

Animal donor organs and tissue have been subjects of study since the 1960s, and some tissues, particularly heart valves, have been commonly utilized. Xenotransplant has the potential to address the critical shortage in organ grafts. The science behind xenotransplant trials has advanced considerably and more human clinical trials utilizing porcine xenografts are quickly approaching.


History
The English word "organ" dates back to the twelfth century, referring to any musical instrument. By the late 14th century, the musical term's meaning had narrowed to refer specifically to the keyboard-based instrument. At the same time, a second meaning arose, in reference to a "body part adapted to a certain function".

Plant organs are made from tissue composed of different types of tissue. The three tissue types are ground, vascular, and dermal. When three or more organs are present, it is called an organ system.

The adjective , also , is used for anything pertaining to the internal organs. Historically, viscera of animals were examined by pagan like the or the in order to divine the future by their shape, dimensions or other factors.

(2026). 9780415311298, Routledge.
This practice remains an important ritual in some remote, tribal societies.

The term "visceral" is contrasted with the term "", meaning "of or relating to the wall of a body part, " The two terms are often used in describing a membrane or piece of connective tissue, referring to the opposing sides.


Antiquity
used the word frequently in his philosophy, both to describe the organs of plants or animals (e.g. the roots of a tree, the heart or liver of an animal) because, in ancient Greek, the word 'organon' means 'tool', and Aristotle believed that the organs of the body were tools for us by means of which we can do things. For similar reasons, his logical works, taken as a whole, are referred to as the because logic is a tool for philosophical thinking. Earlier thinkers, such as those who wrote texts in the Hippocratic corpus, generally did not believe that there were organs of the body but only different parts of the body.Gundert 1992: 465. Gundert, Beate. 1992. "Parts and Their Roles in Hippocratic Medicine," Isis 83: 453–65.

Some alchemists (e.g. ) adopted the assignment between the seven vital organs and the seven classical planets as follows:

(2026). 9780099457879, Arrow.
Heart
Brain
Lungs
Kidneys
Gall bladder
Liver
Spleen

Chinese traditional medicine recognizes eleven organs, associated with the five Chinese traditional elements and with yin and yang, as follows:

Woodyinliver
yanggall bladder
Fireyinheart
yangsmall intestine /
Earthyinspleen
yangstomach
Metalyinlungs
yanglarge intestine
Wateryinkidneys
yangbladder

The Chinese associated the five elements with the five planets (Jupiter, Mars, Venus, Saturn, and Mercury) similar to the way the classical planets were associated with different metals. The yin and yang distinction approximates the modern notion of solid and hollow organs.


See also


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