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Neuroscience is the science of the nervous system (the brain, spinal cord, and peripheral nervous system), its functions, and its disorders. It is a multidisciplinary science that combines physiology, anatomy, molecular biology, developmental biology, cytology, psychology, physics, computer science, chemistry, medicine, statistics, and mathematical modeling to understand the fundamental and emergent properties of , glia, and . The understanding of the biological basis of learning, memory, behavior, perception, and consciousness has been described by Eric Kandel as the "epic challenge" of the Biology.
The scope of neuroscience has broadened over time to include different approaches used to study the nervous system at different scales. The techniques used by have expanded enormously, from molecular and cell biology studies of individual neurons to neuroimaging of Sensory neuron, Motor neuron, and Cognition tasks in the brain.
Early views on the function of the brain regarded it to be a "cranial stuffing" of sorts. In Egypt, from the late Middle Kingdom onwards, the brain was regularly removed in preparation for Mummy. It was believed at the time that the heart was the seat of intelligence. According to Herodotus, the first step of mummification was to "take a crooked piece of iron, and with it draw out the brain through the nostrils, thus getting rid of a portion, while the skull is cleared of the rest by rinsing with drugs."
The view that the heart was the source of consciousness was not challenged until the time of the Greek physician Hippocrates. He believed that the brain was not only involved with sensation—since most specialized organs (e.g., eyes, ears, tongue) are located in the head near the brain—but was also the seat of intelligence. Plato also speculated that the brain was the seat of the rational part of the soul. Aristotle, however, believed the heart was the center of intelligence and that the brain regulated the amount of heat from the heart. (2025). 9780195146943, Oxford University Press, USA. ISBN 9780195146943 This view was generally accepted until the Roman physician Galen, a follower of Hippocrates and physician to Gladiator, observed that his patients lost their mental faculties when they had sustained damage to their brains.
Abulcasis, Averroes, Avicenna, Ibn Zuhr, and Maimonides, active in the Medieval Muslim world, described a number of medical problems related to the brain. In Renaissance, Vesalius (1514–1564), René Descartes (1596–1650), Thomas Willis (1621–1675) and Jan Swammerdam (1637–1680) also made several contributions to neuroscience.
Luigi Galvani's pioneering work in the late 1700s set the stage for studying the electrical excitability of muscles and neurons. In 1843 Emil du Bois-Reymond demonstrated the electrical nature of the nerve signal, whose speed Hermann von Helmholtz proceeded to measure, (2025). 9783319130682, Springer International Publishing. ISBN 9783319130682 and in 1875 Richard Caton found electrical phenomena in the cerebral hemispheres of rabbits and monkeys. Adolf Beck published in 1890 similar observations of spontaneous electrical activity of the brain of rabbits and dogs. Studies of the brain became more sophisticated after the invention of the microscope and the development of a staining by Camillo Golgi during the late 1890s. The procedure used a silver chromate salt to reveal the intricate structures of individual . His technique was used by Santiago Ramón y Cajal and led to the formation of the neuron doctrine, the hypothesis that the functional unit of the brain is the neuron. Golgi and Ramón y Cajal shared the Nobel Prize in Physiology or Medicine in 1906 for their extensive observations, descriptions, and categorizations of neurons throughout the brain.
In parallel with this research, in 1815 Jean Pierre Flourens induced localized lesions of the brain in living animals to observe their effects on motricity, sensibility and behavior. Work with brain-damaged patients by Marc Dax in 1836 and Paul Broca in 1865 suggested that certain regions of the brain were responsible for certain functions. At the time, these findings were seen as a confirmation of Franz Joseph Gall's theory that language was localized and that certain mental process were localized in specific areas of the cerebral cortex. The localization of function hypothesis was supported by observations of epilepsy patients conducted by John Hughlings Jackson, who correctly inferred the organization of the motor cortex by watching the progression of seizures through the body. Carl Wernicke further developed the theory of the specialization of specific brain structures in language comprehension and production. Modern research through neuroimaging techniques, still uses the Brodmann areas cerebral cytoarchitectonic map (referring to the study of Organelle) anatomical definitions from this era in continuing to show that distinct areas of the cortex are activated in the execution of specific tasks.
During the 20th century, neuroscience began to be recognized as a distinct academic discipline in its own right, rather than as studies of the nervous system within other disciplines. Eric Kandel and collaborators have cited David Rioch, Francis O. Schmitt, and Stephen Kuffler as having played critical roles in establishing the field. Rioch originated the integration of basic anatomical and physiological research with clinical psychiatry at the Walter Reed Army Institute of Research, starting in the 1950s. During the same period, Schmitt established a neuroscience research program within the Biology Department at the Massachusetts Institute of Technology, bringing together biology, chemistry, physics, and mathematics. The first freestanding neuroscience department (then called Psychobiology) was founded in 1964 at the University of California, Irvine by James McGaugh. This was followed by the Department of Neurobiology at Harvard Medical School, which was founded in 1966 by Stephen Kuffler.
In the process of treating epilepsy, Wilder Penfield produced maps of the location of various functions (motor, sensory, memory, vision) in the brain. Wilder Penfield redrew the map of the brain — by opening the heads of living patients He summarized his findings in a 1950 book called The Cerebral Cortex of Man. Wilder Penfield and his co-investigators Edwin Boldrey and Theodore Rasmussen are considered to be the originators of the cortical homunculus.
The understanding of neurons and of nervous system function became increasingly precise and molecular during the 20th century. For example, in 1952, Alan Lloyd Hodgkin and Andrew Huxley presented a mathematical model for the transmission of electrical signals in neurons of the giant axon of a squid, which they called "action potentials", and how they are initiated and propagated, known as the Hodgkin–Huxley model. In 1961–1962, Richard FitzHugh and J. Nagumo simplified Hodgkin–Huxley, in what is called the FitzHugh–Nagumo model. In 1962, Bernard Katz modeled neurotransmission across the space between neurons known as synapses. Beginning in 1966, Eric Kandel and collaborators examined biochemical changes in neurons associated with learning and memory storage in Aplysia. In 1981 Catherine Morris and Harold Lecar combined these models in the Morris–Lecar model. Such increasingly quantitative work gave rise to numerous biological neuron models and models of neural computation.
As a result of the increasing interest about the nervous system, several prominent neuroscience organizations have been formed to provide a forum to all neuroscientists during the 20th century. For example, the International Brain Research Organization was founded in 1961, the International Society for Neurochemistry in 1963, the European Brain and Behaviour Society in 1968, and the Society for Neuroscience in 1969. Recently, the application of neuroscience research results has also given rise to Applied science as neuroeconomics, neuroeducation,Zull, J. (2002). The art of changing the brain: Enriching the practice of teaching by exploring the biology of learning. Sterling, Virginia: Stylus Publishing, LLC neuroethics, and neurolaw.
Over time, brain research has gone through philosophical, experimental, and theoretical phases, with work on neural implants and brain simulation predicted to be important in the future.
For example, it has become possible to understand, in much detail, the complex processes occurring within a single neuron. Neurons are cells specialized for communication. They are able to communicate with neurons and other cell types through specialized junctions called , at which electrical or electrochemical signals can be transmitted from one cell to another. Many neurons extrude a long thin filament of axoplasm called an axon, which may extend to distant parts of the body and are capable of rapidly carrying electrical signals, influencing the activity of other neurons, muscles, or glands at their termination points. A nervous system emerges from the assemblage of neurons that are connected to each other in , and networks.
The vertebrate nervous system can be split into two parts: the central nervous system (defined as the brain and spinal cord), and the peripheral nervous system. In many species—including all vertebrates—the nervous system is the most Complex system in the body, with most of the complexity residing in the brain. The human brain alone contains around one hundred billion neurons and one hundred trillion synapses; it consists of thousands of distinguishable substructures, connected to each other in synaptic networks whose intricacies have only begun to be unraveled. At least one out of three of the approximately 20,000 genes belonging to the human genome is expressed mainly in the brain.
Due to the high degree of neuroplasticity of the human brain, the structure of its synapses and their resulting functions change throughout life.
Making sense of the nervous system's dynamic complexity is a formidable research challenge. Ultimately, neuroscientists would like to understand every aspect of the nervous system, including how it works, how it develops, how it malfunctions, and how it can be altered or repaired. Analysis of the nervous system is therefore performed at multiple levels, ranging from the molecular and cellular levels to the systems and cognitive levels. The specific topics that form the main focus of research change over time, driven by an ever-expanding base of knowledge and the availability of increasingly sophisticated technical methods. Improvements in technology have been the primary drivers of progress. Developments in electron microscopy, computer science, electronics, functional neuroimaging, and genetics and genomics have all been major drivers of progress.
Advances in the classification of have been enabled by electrophysiological recording, single-cell genetic sequencing, and high-quality microscopy, which have combined into a single method pipeline called patch-sequencing in which all three methods are simultaneously applied using miniature tools. The efficiency of this method and the large amounts of data that is generated has allowed researchers to make some general conclusions about cell types; for example that the human and mouse brain have different versions of fundamentally the same cell types.
Questions addressed in cellular neuroscience include the mechanisms of how neurons process cell signaling physiologically and electrochemically. These questions include how signals are processed by neurites and somas and how and electrical signals are used to process information in a neuron. Neurites are thin extensions from a neuronal Perikaryon, consisting of (specialized to receive synaptic inputs from other neurons) and (specialized to conduct nerve impulses called ). Somas are the cell bodies of the neurons and contain the nucleus.
Another major area of cellular neuroscience is the investigation of the development of the nervous system. Questions include the patterning and regionalization of the nervous system, axonal and dendritic development, Growth factor, Synaptogenesis and the implication of in neural stem cells, differentiation of neurons and glia (neurogenesis and gliogenesis), and neuronal migration.
Computational neurogenetic modeling is concerned with the development of dynamic neuronal models for modeling brain functions with respect to genes and dynamic interactions between genes, on the cellular level (Computational Neurogenetic Modeling (CNGM) can also be used to model neural systems).
Questions in systems neuroscience include how are formed and used anatomically and physiologically to produce functions such as , multisensory integration, motor coordination, , emotion, learning, and memory. In other words, this area of research studies how connections are made and morphed in the brain, and the effect it has on human sensation, movement, attention, inhibitory control, decision-making, reasoning, memory formation, reward, and emotion regulation.
Specific areas of interest for the field include observations of how the structure of neural circuits effect skill acquisition, how specialized regions of the brain develop and change (neuroplasticity), and the development of brain atlases, or wiring diagrams of individual developing brains.
The related fields of neuroethology and neuropsychology address the question of how neural substrates underlie specific Ethology and human behavior behaviors. Neuroendocrinology and psychoneuroimmunology examine interactions between the nervous system and the endocrinology and immunology systems, respectively. Despite many advancements, the way that networks of neurons perform complex cognition and behaviors is still poorly understood.
Neuroscience is also allied with the social science and behavioral sciences, as well as with nascent interdisciplinary fields. Examples of such alliances include neuroeconomics, decision theory, social neuroscience, and neuromarketing to address complex questions about interactions of the brain with its environment. A study into consumer responses for example uses EEG to investigate neural correlates associated with narrative transportation into stories about energy efficiency.
Neurology works with diseases of the central and peripheral nervous systems, such as amyotrophic lateral sclerosis (ALS) and stroke, and their medical treatment. Psychiatry focuses on affective, behavioral, cognition, and perception disorders. Anesthesiology focuses on perception of pain, and pharmacologic alteration of consciousness. Neuropathology focuses upon the classification and underlying pathogenic mechanisms of central and peripheral nervous system and muscle diseases, with an emphasis on morphologic, microscopic, and chemically observable alterations. Neurosurgery and psychosurgery work primarily with surgical treatment of diseases of the central and peripheral nervous systems.
Neuroscience underlies the development of various neurotherapy methods to treat diseases of the nervous system.IEEE Brain (2019). "Neurotherapy: Treating Disorders by Retraining the Brain". The Future Neural Therapeutics White Paper
Integrative neuroscience describes the effort to combine models and information from multiple levels of research to develop a coherent model of the nervous system. For example, brain imaging coupled with physiological numerical models and theories of fundamental mechanisms may shed light on psychiatric disorders.
Another important area of translational research is brain–computer interfaces (BCIs), or machines that are able to communicate and influence the brain. They are currently being researched for their potential to repair neural systems and restore certain cognitive functions. However, some ethical considerations have to be dealt with before they are accepted.
Other major organizations devoted to neuroscience include the International Brain Research Organization (IBRO), which holds its meetings in a country from a different part of the world each year, and the Federation of European Neuroscience Societies (FENS), which holds a meeting in a different European city every two years. FENS comprises a set of 32 national-level organizations, including the British Neuroscience Association, the German Neuroscience Society (Neurowissenschaftliche Gesellschaft), and the French Société des Neurosciences. The first National Honor Society in Neuroscience, Nu Rho Psi, was founded in 2006. Numerous youth neuroscience societies which support undergraduates, graduates and early career researchers also exist, such as Simply Neuroscience and Project Encephalon.
In 2013, the BRAIN Initiative was announced in the US. The International Brain Initiative was created in 2017, currently integrated by more than seven national-level brain research initiatives (US, Europe, Allen Institute, Japan, China, Australia, Canada, Korea, and Israel) spanning four continents.
Neuroscience educators formed a Faculty for Undergraduate Neuroscience (FUN) in 1992 to share best practices and provide travel awards for undergraduates presenting at Society for Neuroscience meetings.
Neuroscientists have also collaborated with other education experts to study and refine educational techniques to optimize learning among students, an emerging field called educational neuroscience. Federal agencies in the United States, such as the National Institute of Health (NIH) and National Science Foundation (NSF), have also funded research that pertains to best practices in teaching and learning of neuroscience concepts.
One example of such a computer is the SpiNNaker supercomputer.
Sensors can also be made smart with neuromorphic technology. An example of this is the Event Camera's BrainScaleS (brain-inspired Multiscale Computation in Neuromorphic Hybrid Systems), a hybrid analog neuromorphic supercomputer located at Heidelberg University in Germany. It was developed as part of the Human Brain Project's neuromorphic computing platform and is the complement to the SpiNNaker supercomputer, which is based on digital technology. The architecture used in BrainScaleS mimics biological neurons and their connections on a physical level; additionally, since the components are made of silicon, these model neurons operate on average 864 times (24 hours of real time is 100 seconds in the machine simulation) that of their biological counterparts.
Recent advances in neuromorphic microchip technology have led a group of scientists to create an artificial neuron that can replace real neurons in diseases.
Val Danilov I (2023). "The Origin of Natural Neurostimulation: A Narrative Review of Noninvasive Brain Stimulation Techniques." OBM Neurobiology
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Translational research
Recently, the boundaries between various specialties have blurred, as they are all influenced by basic research in neuroscience. For example, brain imaging enables objective biological insight into mental illnesses, which can lead to faster diagnosis, more accurate prognosis, and improved monitoring of patient progress over time.
Major branches
Modern neuroscience education and research activities can be very roughly categorized into the following major branches, based on the subject and scale of the system in examination as well as distinct experimental or curricular approaches. Individual neuroscientists, however, often work on questions that span several distinct subfields.
+ List of the major branches of neuroscience Affective neuroscience Affective neuroscience is the study of the neural mechanisms involved in emotion, typically through experimentation on animal models. Behavioral neuroscience Behavioral neuroscience (also known as biological psychology, physiological psychology, biopsychology, or psychobiology) is the application of the principles of biology to the study of genetic, physiological, and developmental mechanisms of behavior in humans and non-human animals. Cellular neuroscience Cellular neuroscience is the study of neurons at a cellular level including morphology and physiological properties. Clinical neuroscience The scientific study of the biological mechanisms that underlie the disorders and diseases of the nervous system. Cognitive neuroscience Cognitive neuroscience is the study of the biological mechanisms underlying cognition. Computational neuroscience Computational neuroscience is the theoretical study of the nervous system. Cultural neuroscience Cultural neuroscience is the study of how cultural values, practices and beliefs shape and are shaped by the mind, brain and genes across multiple timescales.Chiao, J.Y. & Ambady, N. (2007). Cultural neuroscience: Parsing universality and diversity across levels of analysis. In Kitayama, S. and Cohen, D. (Eds.) Handbook of Cultural Psychology, Guilford Press, New York, pp. 237–254. Developmental neuroscience Developmental neuroscience studies the processes that generate, shape, and reshape the nervous system and seeks to describe the cellular basis of neural development to address underlying mechanisms. Evolutionary neuroscience Evolutionary neuroscience studies the evolution of nervous systems.Eryomin A.L. (2022) Biophysics of Evolution of Intellectual Systems // Biophysics, Vol. 67, No. 2, pp. 320–326. Molecular neuroscience Molecular neuroscience studies the nervous system with molecular biology, molecular genetics, protein chemistry, and related methodologies.Longstaff, Alan; Revest, Patricia (1998). Molecular Neuroscience. Garland Science. ISBN 978-1859962503. Nanoneuroscience An interdisciplinary field that integrates nanotechnology and neuroscience. Neural engineering Neural engineering uses engineering techniques to interact with, understand, repair, replace, or enhance neural systems. Neuroanatomy Neuroanatomy is the study of the anatomy of . (2025). 9780387775791 ISBN 9780387775791Neurochemistry Neurochemistry is the study of how interact and influence the function of neurons. Neuroethology Neuroethology is the study of the neural basis of non-human animals behavior. Neurogastronomy Neurogastronomy is the study of flavor and how it affects sensation, cognition, and memory. (2013). 9780231159111, Columbia University Press. ISBN 9780231159111Neurogenetics Neurogenetics is the study of the genetical basis of the development and function of the nervous system. Neuroimaging Neuroimaging includes the use of various techniques to either directly or indirectly image the structure and function of the brain. Neuroimmunology Neuroimmunology is concerned with the interactions between the nervous and the immune system. Neuroinformatics Neuroinformatics is a discipline within bioinformatics that conducts the organization of neuroscience data and application of computational models and analytical tools. Neurolinguistics Neurolinguistics is the study of the neural mechanisms in the human brain that control the comprehension, production, and acquisition of language. Neuro-ophthalmology Neuro-ophthalmology is an academically oriented subspecialty that merges the fields of neurology and ophthalmology, often dealing with complex systemic diseases that have manifestations in the visual system. Neurophysics Neurophysics is the branch of biophysics dealing with the development and use of physical methods to gain information about the nervous system. Neurophysiology Neurophysiology is the study of the structure and function of the nervous system, generally using physiological techniques that include measurement and stimulation with electrodes or optically with ion- or voltage-sensitive dyes or light-sensitive channels. (2025). 9781402082641 ISBN 9781402082641Neuropsychology Neuropsychology is a discipline that resides under the umbrellas of both psychology and neuroscience, and is involved in activities in the arenas of both basic science and applied science. In psychology, it is most closely associated with biopsychology, clinical psychology, cognitive psychology, and developmental psychology. In neuroscience, it is most closely associated with the cognitive, behavioral, social, and affective neuroscience areas. In the applied and medical domain, it is related to neurology and psychiatry.Gluck, Mark A.; Mercado, Eduardo; Myers, Catherine E. (2016). Learning and Memory: From Brain to Behavior. New York/NY, USA: Worth Publishers. p. 57. ISBN 978-1-319-15405-9. Neuropsychopharmacology Neuropsychopharmacology, an interdisciplinary science related to psychopharmacology and fundamental neuroscience, is the study of the neural mechanisms that drugs act upon to influence behavior. Optogenetics Optogenetics is a biological technique to control the activity of neurons or other cell types with light. Paleoneurobiology Paleoneurobiology is a field that combines techniques used in paleontology and archeology to study brain evolution, especially that of the human brain. Social neuroscience Social neuroscience is an interdisciplinary field devoted to understanding how biological systems implement social processes and behavior, and to using biological concepts and methods to inform and refine theories of social processes and behavior. Systems neuroscience Systems neuroscience is the study of the function of neural circuits and systems.
Careers in neuroscience
The career options for neuroscience graduates vary widely depending on the level of education. At the bachelor’s level, graduates often enter laboratory research, healthcare support, biotechnology, or science communication, though some pursue broader fields such as policy or nonprofit work. With a master’s degree, training may prepare individuals for applied health professions (e.g., occupational therapy, medicine -neurology, psychiatry, neuroimaging-, genetic counseling), research management, or public health. An advanced degree (PhD or equivalent) is usually required for independent research or university teaching.
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Bachelor's Level
Pharmaceutical Sales Assisted living Laboratory Technician Regulatory Affairs Specialist Psychometrist* Medical Technician* Science Writer Clinical Research Assistant Science Advocacy Special Education Assistant Nonprofit Work Patient Care Assistant* Health Educator Orthotic and Prosthetic Technician* EEG Technologist* Lab Animal Care Technician Medical and Healthcare Manager Sales Engineer Forensic Science Technician Law Enforcement Pharmacy Technician* Natural Sciences Manager Public Policy Advertising/Marketing
Master's Level
Nurse Practitioner Neuroimaging Technician Physician's Assistant Teacher Genetic Counselor Epidemiology Occupational Therapist Biostatistician Orthotist/Prosthetist Speech-Language Pathologist Neural Engineer Public Health
Advanced Degree
Medicine (MD, DO) Sensory analysis Research Scientist Pharmacist Dentist Veterinarian Physical Therapist Audiologist Optometrist Lawyer Clinical Psychologist Professor Neuropsychologist Chiropractor
Neuroscience organizations
The largest professional neuroscience organization is the Society for Neuroscience (SFN), which is based in the United States but includes many members from other countries. Since its founding in 1969 the SFN has grown steadily: as of 2010 it recorded 40,290 members from 83 countries. Annual meetings, held each year in a different American city, draw attendance from researchers, postdoctoral fellows, graduate students, and undergraduates, as well as educational institutions, funding agencies, publishers, and hundreds of businesses that supply products used in research.
Public education and outreach
In addition to conducting traditional research in laboratory settings, neuroscientists have also been involved in the promotion of awareness and knowledge about the nervous system among the general public and government officials. Such promotions have been done by both individual neuroscientists and large organizations. For example, individual neuroscientists have promoted neuroscience education among young students by organizing the International Brain Bee, which is an academic competition for high school or secondary school students worldwide. In the United States, large organizations such as the Society for Neuroscience have promoted neuroscience education by developing a primer called Brain Facts, collaborating with public school teachers to develop Neuroscience Core Concepts for K-12 teachers and students, and cosponsoring a campaign with the Dana Foundation called Brain Awareness Week to increase public awareness about the progress and benefits of brain research. In Canada, the Canadian Institutes of Health Research's (CIHR) Canadian National Brain Bee is held annually at McMaster University.
Engineering applications of neuroscience
Neuromorphic computer chips
Neuromorphic engineering is a branch of neuroscience that deals with creating functional of neurons for the purposes of useful computation. The emergent computational properties of neuromorphic computers are fundamentally different from conventional computers in the sense that they are complex system, and that the computational components are interrelated with no central processor.
Nobel prizes related to neuroscience
1904 Physiology Ivan Pavlov 1849–1936 Russian Empire "in recognition of his work on the physiology of digestion, through which knowledge on vital aspects of the subject has been transformed and enlarged" 1906 Physiology Camillo Golgi 1843–1926 Kingdom of Italy "in recognition of their work on the structure of the nervous system" Santiago Ramón y Cajal 1852–1934 Restoration (Spain) 1911 Physiology Allvar Gullstrand 1862– 1930 Sweden "for his work on the dioptrics of the eye" 1914 Physiology Robert Bárány 1876–1936 Austria-Hungary "for his work on the physiology and pathology of the vestibular apparatus" 1932 Physiology Charles Scott Sherrington 1857–1952 United Kingdom "for their discoveries regarding the functions of neurons" Edgar Douglas Adrian 1889–1977 United Kingdom 1936 Physiology Henry Hallett Dale 1875–1968 United Kingdom "for their discoveries relating to chemical transmission of nerve impulses" Otto Loewi 1873–1961 Austria
Germany1938 Physiology Corneille Jean François Heymans 1892–1968 Belgium "for the discovery of the role played by the sinus and aorta in the regulation of respiration" 1944 Physiology Joseph Erlanger 1874–1965 United States "for their discoveries relating to the highly differentiated functions of single nerve fibres" Herbert Spencer Gasser 1888–1963 United States 1949 Physiology Walter Rudolf Hess 1881–1973 Switzerland "for his discovery of the functional organization of the interbrain as a coordinator of the activities of the internal organs" António Caetano Egas Moniz 1874–1955 Portugal "for his discovery of the therapeutic value of leucotomy in certain psychoses" 1955 Chemistry Vincent du Vigneaud 1901–1978 United States "for his work on biochemically important sulphur compounds, especially for the first synthesis of a Oxytocin" (Oxytocin) 1957 Physiology Daniel Bovet 1907–1992 Italy "for his discoveries relating to synthetic compounds that inhibit the action of certain body substances, and especially their action on the vascular system and the skeletal muscles" 1961 Physiology Georg von Békésy 1899–1972 United States "for his discoveries of the physical mechanism of stimulation within the cochlea" 1963 Physiology John Carew Eccles 1903–1997 Australia "for their discoveries concerning the ionic mechanisms involved in excitation and inhibition in the peripheral and central portions of the nerve cell membrane" Alan Lloyd Hodgkin 1914–1998 United Kingdom Andrew Huxley 1917–2012 United Kingdom 1967 Physiology Ragnar Granit 1900–1991 Finland
Sweden"for their discoveries concerning the primary physiological and chemical visual processes in the eye" Haldan Keffer Hartline 1903–1983 United States George Wald 1906–1997 United States 1970 Physiology Julius Axelrod 1912–2004 United States "for their discoveries concerning the humoral Neurotransmitter and the mechanism for their storage, release and inactivation" Ulf von Euler 1905–1983 Sweden Bernard Katz 1911–2003 United Kingdom 1973 Physiology Karl von Frisch 1886–1982 Austria "for their discoveries concerning organization and elicitation of individual and social behaviour patterns" Konrad Lorenz 1903–1989 Austria Nikolaas Tinbergen 1907–1988 Netherlands 1977 Physiology Roger Guillemin 1924–2024 France "for their discoveries concerning the peptide hormone production of the brain" Andrew Schally 1926–2024 Poland
1981 Physiology Roger W. Sperry 1913–1994 United States "for his discoveries concerning the functional specialization of the cerebral hemispheres" David H. Hubel 1926–2013 Canada "for their discoveries concerning information processing in the visual system" Torsten Wiesel 1924– Sweden 1986 Physiology Stanley Cohen 1922–2020 United States "for their discoveries of " Rita Levi-Montalcini 1909–2012 Italy 1991 Physiology Erwin Neher 1944– Germany "For their discoveries concerning the function of single ion channels in cells" Bert Sakmann 1942– Germany 1997 Physiology Stanley B. Prusiner 1942– United States "for his discovery of - a new biological principle of infection" 1997 Chemistry Jens C. Skou 1918–2018 Denmark "for the first discovery of an ion-transporting enzyme, Na+, K+ -ATPase" 2000 Physiology Arvid Carlsson 1923–2018 Sweden "for their discoveries concerning signal transduction in the nervous system" Paul Greengard 1925–2019 United States Eric Kandel 1929– United States 2003 Chemistry Roderick MacKinnon 1956– United States "for discoveries concerning channels in cell membranes ... for structural and mechanistic studies of ion channels" 2004 Physiology Richard Axel 1946– United States "for their discoveries of and the organization of the olfactory system" Linda B. Buck 1947– United States 2012 Chemistry Robert Lefkowitz 1943– United States "for studies of G-protein-coupled receptors"" Brian Kobilka 1955– United States 2014 Physiology John O'Keefe 1939– United States
United Kingdom"for their discoveries of Place cell and Grid cell cells that constitute a positioning system in the brain" May-Britt Moser 1963– Norway Edvard I. Moser 1962– Norway 2017 Physiology Jeffrey C. Hall 1939– United States "for their discoveries of molecular mechanisms controlling the circadian rhythm" Michael Rosbash 1944– United States Michael W. Young 1949– United States 2021 Physiology David Julius 1955– United States "for their discoveries of receptors for temperature and touch" Ardem Patapoutian 1967– Lebanon
United States 2024 Physics John Hopfield 1933– United States "for foundational discoveries and inventions that enable machine learning with artificial neural networks" Geoffrey Hinton 1947– United Kingdom
See also
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