Llama

 ( Camelids ) 

The following characteristics apply especially to llamas. Dentition of adults: incisors canines , premolars , molars ; total 32. In the upper jaw, a compressed, sharp, pointed laniariform incisor near the hinder edge of the premaxilla is followed in the male at least by a moderate-sized, pointed, curved true canine tooth in the anterior part of the maxilla. The isolated canine-like premolar that follows in the camels is not present. The teeth of the molar series, which are in contact with each other, consist of two very small premolars (the first almost rudimentary) and three broad molars, generally constructed like those of Camelus. In the lower jaw, the three incisors are long, spatulate, and procumbent; the outer ones are the smallest. Next to these is a curved, suberect canine, followed after an interval by an isolated minute and often deciduous simple conical premolar; then a contiguous series of one premolar and three molars, which differ from those of Camelus in having a small accessory column at the anterior outer edge.

The skull generally resembles that of Camelus, with a larger brain cavity and orbits and less-developed cranial ridges due to its smaller size. The nasal bones are shorter and broader and are joined by the premaxilla.

Vertebrae:

• cervical 7,
• dorsal 12,
• lumbar 7,
• sacral 4,
• caudal 15 to 20.

The ears are rather long and slightly curved inward, characteristically known as "banana" shaped. There is no dorsal hump. The feet are narrow, the toes being more separated than in the camels, each having a distinct plantar pad. The tail is short, and the fiber is long, woolly, and soft.

In essential structural characteristics, as well as in general appearance and habits, all the animals of this genus very closely resemble each other, so whether they should be considered as belonging to one, two, or more species is a matter of controversy among natural history.

The question is complicated by the circumstances of most individuals who have come under observation, either in a completely or partially domesticated state. Many are also descended from ancestors previously domesticated, a state that tends to produce a certain amount of variation from the original type. The four forms commonly distinguished by the inhabitants of South America are recognized as distinct species, though there are difficulties in defining their distinctive characteristics.

These are:

• the llama, Lama glama (Carl Linnaeus);
• the alpaca, Lama pacos (Linnaeus);
• the guanaco (from the Quechua huanaco), Lama guanicoe (Müller); and
• the vicuña, Lama vicugna (Molina)

The llama and alpaca are only known in the domestic state and are variable in size and of many colors, often white, brown, or piebald. Some are grey or black. The guanaco and vicuña are wild. The guanaco is endangered; it has a nearly uniform light-brown color, passing into white below.

The guanaco and vicuña certainly differ: The vicuña is more petite, more slender in its proportions, and has a shorter head than the guanaco.

The vicuña lives in on the tierra helada bordering the janca, amidst rocks and precipices, occurring in various suitable localities throughout Peru, in the southern part of Ecuador, and as far south as the middle of Bolivia. Its manners very much resemble those of the chamois of the European Alps; it is as vigilant, wild, and timid.

Vicuña fiber is extremely delicate and soft and highly valued for weaving, but the quantity that each animal produces is small. Alpacas are primarily descended from wild vicuña ancestors. In contrast, domesticated llamas are descended primarily from wild guanaco ancestors, although a considerable amount of hybridization between the two species has occurred.

Differential characteristics between llamas and alpacas include the llama's larger size, longer head, and curved ears. Alpaca fiber is generally more expensive but not always more valuable. Alpacas tend to have a more consistent color throughout the body. The most apparent visual difference between llamas and is that camels have a humps and llamas do not.

Llamas are not , pseudo-ruminants, or modified ruminants. They do have a complex three-compartment stomach that allows them to digest lower quality, high cellulose foods. The stomach compartments allow for fermentation of tricky foodstuffs, followed by regurgitation and re-chewing. (cows, sheep, goats) have four compartments, whereas llamas have only three stomach compartments: the rumen, omasum, and abomasum.

In addition, the llama (and other camelids) have an extremely long and complex large intestine (colon). The large intestine's role in digestion is to reabsorb water, vitamins, and electrolytes from food waste passing through it. The length of the llama's colon allows it to survive on much less water than other animals. This is a major advantage in arid climates where they live.

Like humans, llama males and females mature sexually at different rates. Females reach puberty at about 12 months old; males do not become sexually mature until around three years of age.

For field mating, a female is turned into a field with a male llama and left there for some time. This is the easiest method in terms of labor but the least useful in predicting a likely birth date. An ultrasound test can be performed, and together with the exposure dates, a better idea of when the cria is expected can be determined.

Hand mating is the most efficient method, but it requires the most work on the part of the human involved. A male and female llama are put into the same pen, and mating is monitored. They are then separated and re-mated every other day until one refuses the mating. Usually, one can get in two matings using this method, though some stud males routinely refuse to mate a female more than once. The separation presumably helps to keep the sperm count high for each mating and also helps to keep the condition of the female llama's reproductive tract more sound. If the mating is unsuccessful within two to three weeks, the female is mated again.

Llamas are now utilized as certified therapy animals in nursing homes and hospitals. Rojo the Llama, located in the Pacific Northwest was certified in 2008. The Mayo Clinic says animal-assisted therapy can reduce pain, depression, anxiety, and fatigue. This type of therapy is growing in popularity, and several organizations throughout the United States participate.

When correctly reared, llamas spitting at a human is a rare thing. Llamas are very social herd animals, however, and sometimes spit at each other to discipline lower-ranked llamas. A llama's social rank in a herd is never static. They can always move up or down the social ladder by picking small fights. This is usually done between males to see which will become dominant. Their fights are visually dramatic, characterized by spitting, ramming each other with their chests, neck wrestling, and kicking, mainly to knock the other off balance. The females are usually only seen spitting to control other herd members. One may determine how agitated the llama is by the materials in the spit. The more irritated the llama is, the further back into each of the three stomach compartments it will try to draw materials from for its spit.

While the social structure might constantly change, they live as a family and care for each other. If one notices a strange noise or feels threatened, an alarm call - a loud, shrill sound that rhythmically rises and falls - is sent out, and all others become alert. They will often hum to each other as a form of communication.

The llama's groaning noises or going "mwa" (/mwaʰ/) is often a sign of fear or anger. Unhappy or agitated llamas will lay their ears back, while ears being perked upwards is a sign of happiness or curiosity.

An "orgle" is the mating sound of a llama or alpaca, made by the sexually aroused male. The sound is reminiscent of gargling but with a more forceful, buzzing edge. Males begin the sound when they become aroused and continue throughout copulation.

Research suggests using multiple guard llamas is not as effective as one. Multiple males tend to bond with one another rather than with the livestock and may ignore the flock. A gelded male of two years of age bonds closely with its new charges and is instinctively very effective in preventing predation. Some llamas bond more quickly to sheep or goats if introduced just before lambing. Many sheep and goat producers indicate a special bond quickly develops between lambs and their guard llama, and the llama is particularly protective of the lambs.

Using llamas as guards has reduced the losses to predators for many producers. The value of the livestock saved each year exceeds a llama's purchase cost and annual maintenance. Although not every llama is suited to the job, most are a viable, nonlethal alternative for reducing predation, requiring no training and little care.

Carl Troll has argued that the large numbers of llamas found in the southern Peruvian highlands were an essential factor in the rise of the Inca Empire.