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A kinglet is a small in the family Regulidae. Species in this family were formerly classified with the Old World warblers. "Regulidae" is derived from the word regulus for "petty king" or prince, and refers to the coloured crowns of adult birds. This family has representatives in and . There are six species in this family; one, the Madeira firecrest ( Regulus madeirensis), was only recently split from the as a separate species. The ruby-crowned kinglet differs sufficiently in its voice and plumage to be afforded its own genus, Corthylio.


Description
Kinglets are among the smallest of all , ranging in size from and weighing ; the sexes are the same size. They have medium-length wings and tails, and small needle-like . The is overall grey-green, offset by pale wingbars, and the tail tip is . Five species have a single stiff feather covering the nostrils, but in the ruby-crowned kinglet this is replaced by several short, stiff bristles. Most kinglets have distinctive head markings, and the males possess a colourful crown patch. In the females, the crown is duller and yellower. The long feathers forming the central crown stripe can be erected; they are inconspicuous most of the time, but are used in courtship and territorial displays when the raised crest is very striking.Martens, Jochen; Päckert, Martin "Family Regulidae (Kinglets & Firecrests)" pp. 330–349 in
(2025). 9788496553064, .

There are two species of different genera in North America with largely overlapping distributions, and two in Eurasia that also have a considerable shared range. In each continent, one species (goldcrest in Eurasia and golden-crowned kinglet in North America) is a conifer specialist; these have deeply grooved pads on their feet for perching on conifer twigs and a long hind toe and claw for clinging vertically. The two generalists, ruby-crowned kinglet and common firecrest, hunt more in flight and have smoother soles, shorter hind claws and a longer tail.


Taxonomy
The kinglets are a small group of birds sometimes included in the Old World warblers but frequently given family status, especially as recent research showed that, despite superficial similarities, the crests are phylogenetically remote from the warblers. The name of the family derives from the regulus, a diminutive of rex, "a king",
(2025). 9780550101853, Chambers.
and refer to the characteristic orange or yellow crests of adult kinglets (aside from the red crest of Corthylio). The kinglets were allocated to the warbler genus by English John Latham in 1790, but moved to their current genus by French in 1800.

Most members of the genus Regulus are similar in size and colour pattern. The exception is the ruby-crowned kinglet, the largest species, which has a strongly red crest and no black crown stripes. It has distinctive vocalisations, and is different enough from the Old World kinglets and the other American species, the golden-crowned kinglet, to be assigned to a separate genus, Corthylio.


Species in taxonomic order
  • Genus Corthylio:
    • Ruby-crowned kinglet ( C. calendula)
  • Genus Regulus:


Distribution and habitat
Kinglets are birds of the and , with representatives in temperate North America, Europe and Asia, northernmost Africa, and the . They are adapted to forests, although there is a certain amount of adaptability and most species will use other habitats, particularly during migration. In Macaronesia, they are adapted to and .


Behaviour

Diet and feeding
The tiny size and rapid metabolism of kinglets means that they must constantly forage in order to provide their energy needs. They will continue feeding even when building. Kinglets prevented from feeding may lose a third of their body weight in twenty minutes and may starve to death in an hour. Kinglets are insectivores, preferentially feeding on prey such as and that have soft cuticles. Prey is generally gleaned from the branches and leaves of trees, although in some circumstances prey may be taken on the wing or from the on the ground.


Life cycle
Kinglet nests are small, very neat cups, almost spherical in shape, made of moss and lichen held together with spiderwebs and hung from twigs near the end of a high branch of a conifer. They are lined with hair and feathers, and a few feathers are placed over the opening. These characteristics provide good insulation against the cold environment. The female lays 7 to 12 eggs, which are white or pale buff, some having fine dark brown spots. Because the nest is small, they are stacked in layers. The female incubates; she pushes her legs (which are well supplied with blood vessels, hence warm) down among the eggs. A unique feature of kinglets is the "size hierarchy" among eggs, with early-laid eggs being smaller than later ones.Haftorn, Svein; "Clutch size, intraclutch egg size variation, and breeding strategy in the Goldcrest Regulus regulus"; in Journal of Ornithology, Volume 127, Number 3 (1986), 291-301.

Eggs hatch asynchronously after 15 to 17 days. The young stay in the nest for 19 to 24 days. After being fed, nestlings make their way down to the bottom of the nest, pushing their still-hungry siblings up to be fed in their turn (but also to be cold).

Kinglets are the most fecund and shortest-living of all altricial birds,Sibly Richard M., Witt, Christopher C., Wright, Natalie A., Venditti, Chris, Jetze, Walter and Brown, James H.; "Energetics, lifestyle, and reproduction in birds" ; in Proceedings of the National Academy of Sciences; 109 (27); pp. 10937-10941 and probably the shortest-lived apart from a few smaller species. Adult mortality for the goldcrest is estimated at over 80 percent per yearIn Ricklefs, R.E.; "Sibling competition, hatching asynchrony, incubation period, and lifespan in altricial birds"; in Power, Dennis M. (editor); Current Ornithology. Vol. 11. and the maximum lifespan is only six years.Wasser, D. E. and Sherman, P.W.; "Avian longevities and their interpretation under evolutionary theories of senescence" in Journal of Zoology 2 November 2009


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