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The cochineal ( , ; Dactylopius coccus) is a in the suborder , from which the natural is derived. A primarily sessile native to tropical and subtropical South America through North America ( and the Southwest ), this insect lives on in the genus , feeding on plant moisture and nutrients. The insects are found on the pads of prickly pear cacti, collected by brushing them off the plants, and dried.

The insect produces that deters predation by other insects. Carminic acid, typically 17–24% of dried insects' weight, can be extracted from the body and eggs, then mixed with aluminium or calcium salts to make carmine dye, also known as cochineal. Today, carmine is primarily used as a and in lipstick ( or ).

Carmine dye was used in the Americas for coloring fabrics and became an important in the 16th century during the colonial period. Production of cochineal is depicted in the (1565). After synthetic pigments and dyes such as were invented in the late 19th century, use of natural-dye products gradually diminished. Fears over the safety of artificial food additives renewed the popularity of cochineal dyes, and the increased demand has made cultivation of the insect profitable again, with being the largest producer, followed by , , and the .

Other species in the genus can be used to produce "cochineal extract", and are extremely difficult to distinguish from D. coccus, even for expert taxonomists; the scientific term D. coccus and the vernacular "cochineal insect" are sometimes used, intentionally or casually, and possibly with misleading effect, to refer to other species.


Etymology
The word cochineal is derived from the French cochenille, derived from Spanish cochinilla, in turn derived from Latin coccinus, from Greek κόκκινος kokkinos, "scarlet" from κόκκος kokkos (Latin equivalent coccum) referring in this case either to the oak berry (actually the insects of the genus Kermes) or to a red dye made from the crushed bodies thereof.. The related in sense word kermes also refers to the source of the red Mediterranean dye also called , which was used in Europe to color cloth red before cochineal was imported from the New World to Spain in the 1520s. Some sources identify the Spanish source word cochinilla as the word for "wood louse", which is a diminutive form of cochino "pig".


Dactylopius coccus

Life cycle
Cochineal insects are soft-bodied, flat, oval-shaped scale insects. The females, wingless and about long, cluster on cactus pads. They penetrate the cactus with their beak-like mouthparts and feed on its juices, remaining immobile unless alarmed. After mating, the fertilised female increases in size and gives birth to tiny nymphs. The nymphs secrete a waxy white substance over their bodies for protection from water loss and excessive sun. This substance makes the cochineal insect appear white or grey from the outside, though the body of the insect and its nymphs produces the red pigment, which makes the insides of the insect look dark purple. Adult males can be distinguished from females in that males have wings, and are much smaller.
(2025). 9780674018273, Belknap Press of Harvard University Press. .

The cochineal disperses in the first nymph stage, called the "crawler" stage. The juveniles move to a feeding spot and produce long wax filaments. Later, they move to the edge of the cactus pad, where the wind catches the wax filaments and carries the insects to a new host. These individuals establish feeding sites on the new host and produce a new generation of cochineals. Male nymphs feed on the cactus until they reach sexual maturity. At this time, they can no longer feed at all and live only long enough to fertilise the eggs. They are, therefore, seldom observed. In addition, females typically outnumber males due to environmental factors.

(2025). 9780520231573, University of California Press.


Host cacti
Dactylopius coccus is native to tropical and subtropical South America and North America in Mexico, where their host cacti grow natively. They have been widely introduced to many regions where their host cacti also grow. About 200 species of Opuntia cacti are known, and while it is possible to cultivate cochineal on almost all of them, the most common is Opuntia ficus-indica. D. coccus has only been noted on Opuntia species, including O. amyclaea, O. atropes, O. cantabrigiensis, O. brasilienis, O. ficus-indica, O. fuliginosa, O. jaliscana, O. leucotricha, O. lindheimeri, O. microdasys, O. megacantha, O. pilifera, O. robusta, O. sarca, O. schikendantzii, O. stricta, O. streptacantha, and O. tomentosa. Feeding cochineals can damage and kill the plant. Other cochineal species feed on many of the same Opuntia, and the wide range of hosts reported for D. coccus likely is because of the difficulty in distinguishing it from other Dactylopius species.
(2025). 9780804716673, Stanford University Press.


Predation
Several natural enemies can reduce the population of the insects on hosts. Of all the predators, insects seem to be the most important group. Insects and their larvae such as (order ), which destroy the cactus, and predators such as (), various (such as and ), (), and ants () have been identified, as well as numerous . Many birds, human-, rodents (especially rats), and reptiles, also prey on cochineal insects.


Farming
A cactus farm for the production of cochineal is traditionally known as a nopalry. The two methods of farming cochineal are traditional and controlled. Cochineals are farmed in the traditional method by planting infected cactus pads or infesting existing cacti with cochineals and harvesting the insects by hand. The controlled method uses small baskets called Zapotec placed on host cacti. The baskets contain clean, fertile females that leave the nests and settle on the cactus to await fertilization by the males. In both cases, the cochineals must be protected from , cold, and rain. The complete cycle lasts three months, during which time the cacti are kept at a constant temperature of 27 °C (81 °F). At the end of the cycle, the new cochineals are left to reproduce or are collected and dried for dye production.

To produce dye from cochineals, the insects are collected when they are around 90 days old. Harvesting the insects is labour-intensive, as they must be individually knocked, brushed, or picked from the cacti and placed into bags. The insects are gathered by small groups of collectors who sell them to local processors or exporters.

In regions dependent on cochineal production, measures are taken seriously. For small-scale cultivation, manual methods of control have proved to be the safest and most effective. For large-scale cultivation, advanced pest control methods have to be developed, including alternative bioinsecticides or traps with .


Failed farming in Australia
Opuntia species, known commonly as prickly pears, were first brought to in an attempt to start a cochineal dye industry in 1788. Captain collected a number of cochineal-infested plants from Brazil on his way to establish the first European settlement at Botany Bay, part of which is now , New South Wales. At that time, Spain and Portugal had a worldwide cochineal dye monopoly via their New World colonial sources, and the British desired a source under their own control, as the dye was important to their clothing and garment industries; it was used to color the British soldiers' red coats, for example. The attempt was a failure in two ways: the Brazilian cochineal insects soon died off, but the cacti thrived, eventually overrunning about of eastern Australia.. The cacti were eventually brought under control in the 1920s by the deliberate introduction of a South American moth, Cactoblastis cactorum, the larvae of which feed on the cactus.


Failed farming in Ethiopia
The nopal pear has been traditionally eaten in parts of northern Ethiopia, where it is utilized more than cultivated. Carmine cochineal was introduced into northern Ethiopia early in the 2000s to be cultivated among farming communities. Foodsafe exported 2000 tons of dried carmine cochineal over 3 years. Https://imbarex.com/< /ref>

A conflict of interest among communities led to closure of the cochineal business in Ethiopia, but the insect spread and became a pest. Cochineal infestation continued to expand after the cochineal business had ended. Control measures were unsuccessful and by 2014 about of cactus land had become infested with cochineal.Tesfay Belay Reda. 2014. Cactus Pear & Carmine Cochineal: introduction & use in Ethiopia. Lambert Academic Publishing.


Biocontrol in South Africa
There has been a population of Dactylopius insects on prickly pear cactuses around in ; several cochineal species were introduced to South Africa, with use encouraged as a biocontrol for different invasive cactus plant species. ISBN 1-86849-168-4


Carmine

Preparation
Cochineal dyes are one of three groups of red insect dyes, all of which are anthraquinone derivatives. The major color components in their respective chemical structures are (in cochineal dyes), (in kermes dye) and (in lac dye).

Carminic acid is extracted from the female cochineal insects and is treated to produce , which can yield shades of red such as and scarlet. The dried body of the female insect is 14–26% carminic acid.

Workers collect the female cochineal insects from their host plants. The insects are killed by immersion in hot water or by exposure to sunlight, steam, or the heat of an oven. Each method produces a different color that results in the varied appearance of commercial cochineal. The insects must be dried to about 30% of their original body weight before they can be stored without decaying. It takes about 70,000 insects to make of cochineal dye.

The two principal forms of cochineal dye are cochineal extract, a coloring made from the raw dried and pulverised bodies of insects, and carmine, a more purified coloring made from the cochineal. To prepare carmine, the powdered insect bodies are boiled in or a solution, the insoluble matter is removed by filtering, and is added to the clear salt solution of carminic acid to precipitate the red salt. Purity of color is ensured by the absence of . Stannous chloride, , , or may be added to regulate the formation of the precipitate. For shades of , lime is added to the alum.


History

Pre-Columbian dye
Traditionally, cochineal was used for coloring . Cochineal dye was used by the and Maya peoples of North and Central America as early as the second century BC.
(2025). 9781473630819, John Murray.
Inhabitants of have been producing cochineal dyes for textiles since early in the Middle Horizon period (600–1000 CE). Cochineal dye was extensively used in the Pre-Columbian era, often for ceremonial textiles and those worn by rulers.

The dye bonds best with animal fibers rather than plant fibers and was most effective for dyeing wool from and other , rabbit fur, and feathers. It was also used on cottons and plant-based fabrics, to less effect. Some examples of early cloth have survived in extremely dry areas in Peru. In addition, the use of cochineal is literally illustrated in drawings on codices and maps. Production of cochineal dyes became well-developed under , and beautiful examples of woven cloth colored by cochineal remain from and .

Cochineal's importance is also indicated by its prominence in tribute lists such as the Matrícula de Tributos. Eleven cities conquered by in the 15th century paid a yearly tribute of 2000 decorated cotton blankets and 40 bags of cochineal dye each.


Use as pigment
Prior to the Spanish invasion, Aztecs also used cochineal pigments in their manuscripts. The 16th century contains a variety of illustrations with multiple variations of the . Specifically in the case of achiotl (light red), technical analysis of the paint reveals multiple layers of the pigment although the layering of the pigment is not visible to the naked eye. Therefore, it proves that the process of applying multiple layers is more significant in comparison to the actual color itself. Furthermore, the process of layering the various hues of the same pigment on top of each other enabled the Aztec artists to create variations in the intensity of the subject matter. A bolder application of pigment draws the viewer's eye to the subject matter which commands attention and suggests a power of the viewer. A weaker application of pigment commands less attention and has less power. This would suggest that the Aztec associated the intensity of pigments with the idea of power and life.

Pigments are insoluble finely ground particles which are mixed with a liquid to make a paint. To be useful as a pigment, a substance should be insoluble in the vehicle with which it is mixed, in contrast to a dye which is soluble. The activity of carmines can vary widely depending on their preparation and composition: they tend to be unstable and can vary in solubility depending on pH.

Recipes for artists' use of crimson appear in many early painting and alchemical handbooks throughout the Middle Ages. Red were known to be particularly unstable as early as the 1400s. When cochineal lakes were introduced in Europe, artists soon found that they were not . The paint turns brown and fades in sunlight, although it is somewhat more permanent if mixed with oil rather than water color. As a result, carmine's use as a pigment was discouraged: its primary use was as a dye rather than in paints.


Comparable colors
In Europe, there was no comparable red dye or pigment. The closest color was Kermes (technically, ), one of the oldest organic pigments. Its key ingredient, kermesic acid, was also extracted from an insect, , which lives on Quercus coccifera oaks native to the Near East, and the European side of the Mediterranean Basin. Kermes was used as a dye and a in ancient Egypt, Greece, Armenia and the Near East.Barber, E. J. W. (1991). Prehistoric Textiles. Princeton University Press.


Colonial export
The Spanish conquest of the Aztec Empire in the 16th century introduced new colors to peoples on both sides of the Atlantic. The Spanish were quick to exploit the vibrant, intense color of cochineal for new trade opportunities. attained great status and value in Europe.

During the colonial period, with the introduction of sheep to Latin America, the use of cochineal increased. It provided the most intense color and it set more firmly on garments compared to clothes made of materials of pre-Hispanic origin such as or and fibers. In general, cochineal is more successful on protein-based animal fibres (including silk) than plant-based material.

Once the European market discovered the qualities of this product ( grana fina), the demand for it increased dramatically.

(2025). 9780816056736, California State University. .
became the region's second-most-valuable export next to silver. The dyestuff was used throughout Europe and was so highly prized, its price was regularly quoted on the London and Amsterdam Commodity Exchanges (with the latter one beginning to record it in 1589). By the 17th century cochineal was a commodity traded as far away as India.

The production and the use of luxury colors and textiles were regulated in countries such as Spain and Italy. Dyestuffs produced from the cochineal insect were used for dyeing the clothes of , , and the .. In 1454, Pope Paul II officially changed the color of the robes worn by Catholic cardinals from "Cardinal's purple" to vibrant red. By 1558, their red robes would have been created with American cochineal. By the 1600s, cochineal also gave the English "Redcoats" their distinctive officers' uniforms. Carmine became strong competition for other colorants such as root, kermes, , Armenian cochineal, , and . It became the most important insect dye used in the production of hand-woven oriental rugs, almost completely displacing lac. It was also used for , and .

Spanish influence also changed the way in which Aztecs used pigments, particularly in their manuscripts. The use of cochineal in manuscripts was replaced by Spanish dyes like minium and .

(2025). 9781606063293, The Getty Research Institute.
The image of Moctezuma's death (seen to the right) uses both indigenous and Spanish pigments, and is therefore representative of the transition and influence between cultures.

During the colonial period in Latin America, many indigenous communities produced cochineal under a type of contract known as Repartimiento de Mercancías. This was a type of "contract forwarding" agreement, in which a trader lent money to producers in advance, with a "" to buy the product once it was harvested. Communities with a history of cochineal production and export have been found to have lower poverty rates and higher female literacy, but also smaller indigenous populations.


Production elsewhere
In 1777, French Nicolas-Joseph Thiéry de Menonville, presenting himself as a botanizing physician, smuggled the insects and pads of the Opuntia cactus to . This particular collection failed to thrive and ultimately died out, leaving the Mexican monopoly intact. After the Mexican War of Independence in 1810–1821, the Mexican monopoly on cochineal came to an end. Large-scale production of cochineal emerged, especially in Guatemala and the Canary Islands; it was also cultivated in Spain and North Africa.


Competition from artificial dyes
The demand for cochineal fell sharply in the middle of the 19th century, with the appearance of artificial dyes such as . This caused a significant financial shock in Spain as a major industry almost ceased to exist. The delicate manual labour required for the breeding of the insect could not compete with the modern methods of the new industry, and even less so with the lowering of production costs. The " tuna blood" dye (from the Mexican name for the Opuntia fruit) stopped being used and trade in cochineal almost totally disappeared in the course of the 20th century. For a time, the breeding of cochineal was done mainly for the purposes of maintaining the tradition rather than to satisfy any sort of demand.

However, the product has become commercially valuable again. One reason for the increasing interest in natural dyes is consumer concern over the possibility that some commercial synthetic red dyes and food colorings may be carcinogenic. Being natural is not a guarantee of safety, but studies show that cochineal is neither carcinogenic nor toxic. Cochineal does, however, have a slight potential to trigger an allergic reaction.


Modern uses
Cochineal continues to be used as a fabric dye, a dye and as a . It is also used in as a preparatory stain for the examination of tissues and carbohydrates.

As of 2005, Peru produced 200 tons of cochineal dye per year and the Canary Islands produced 20 tons per year. Chile and Mexico also export cochineal. France is believed to be the world's largest importer, and and also import the insect. Much of these imports are processed and re-exported to other developed economies. As of 2005, the market price of cochineal was between US$50 and 80 per kilogram, while synthetic raw food dyes are available at prices as low as $10–20 per kilogram.

Natural carmine dye used in food and cosmetics can render the product unacceptable to or consumers. Many consider carmine-containing food forbidden ( ) because the dye is extracted from insects and all insects except the are haram in . also avoid food containing this additive, though it is not , and some authorities allow its use because the insect is dried and reduced to powder.Pischei Teshuvah Yoreh Deah 87-20

Cochineal is one of the few water-soluble colorants to resist degradation with time. It is one of the most light- and heat-stable and oxidation-resistant of all the natural organic colorants and is even more stable than many synthetic food colors. The water-soluble form, cochineal extract, is used in a wide variety of beverages; the insoluble form, carmine, is used in a wide variety of products. They can be found in , , processed products (meat products cannot be colored in the United States unless they are labeled as such), , , alcoholic drinks, bakery products and toppings, , , icings, pie fillings, , preserves, desserts, beverages, varieties of and other products, , and sweets.

Carmine is considered safe enough for cosmetic use in the eye area. A significant proportion of the insoluble carmine pigment produced is used in the cosmetics industry for hair- and skin-care products, , face powders, , and blushes. A bright red dye and the stain carmine used in microbiology is often made from the carmine extract, too. The pharmaceutical industry uses cochineal to color pills and .

Cochineal-colored wool and cotton continue to be important materials for Mexican folk art and crafts.Wood, W. W. (2008). Made in Mexico: Zapotec weavers and the global ethnic art market. Indiana University Press.

(2025). 9781588393616, Metropolitan Museum of Art. .
.
Some towns in the state of continue to follow traditional practices of producing and using cochineal when making handmade textiles. In , Heifer International has partnered with local women who wished to reintroduce traditional artisanal practices of cochineal production and use.

Because it has a complicated structure involving multiple chemical groups, it is very difficult to create a synthetic molecule for cochineal. In 1991, carminic acid was first synthesized in the laboratory by organic chemists. In 2018, researchers genetically engineered the fungus Aspergillus nidulans to produce carminic acid; the bacterium was engineered to produce carminic acid in 2021.


Risks and labeling
In spite of the widespread use of carmine-based dyes in food and cosmetic products, a small number of people have been found to experience occupational asthma, and cosmetic allergies (such as allergic rhinitis and ), -mediated respiratory hypersensitivity, and in rare cases anaphylactic shock.
(2025). 9780851996073, CABI Pub..
In 2009, the FDA ruled that labels of cosmetics and food that include cochineal extract must include that information on their labels (under the name "cochineal extract" or "carmine").FDA. Guidance for Industry: Cochineal Extract and Carmine: Declaration by Name on the Label of All Foods and Cosmetic Products That Contain These Color Additives; Small Entity Compliance Guide . Silver Spring, MD:U.S. Food & Drug Administration (updated June 7, 2011). accessed. In 2006, the FDA stated it found no evidence of a "significant hazard" to the general population. In the EU, authorities list carmine as additive E120 in the . An artificial, non-allergenic cochineal dye is labeled E124.


Explanatory notes

See also
  • Kermes (dye)


Further reading


External links
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