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Biodiversity refers to the variety and variability of Life. It can be measured at multiple levels, including genetic variability, species diversity, ecosystem diversity and Phylogenetics diversity. Diversity is unevenly distributed across the planet and is highest in the tropics, largely due to the region's warm climate and high primary productivity. Although tropical forests cover less than one-fifth of Earth's land surface, they host approximately half of the world's species. Patterns such as the latitudinal gradients in species diversity are observed in both marine and terrestrial organisms.
Since the emergence of Abiogenesis, biodiversity has undergone significant changes, including six major mass extinctions and several smaller events. The Phanerozoic eon (the past 540 million years) saw a rapid expansion of biodiversity, notably during the Cambrian explosion, when many multicellular Phylum first appeared. Over the next 400 million years, biodiversity repeatedly declined due to mass extinction events. These included the Carboniferous rainforest collapse and the Permian–Triassic extinction event 251 million years ago—which caused the most severe biodiversity loss in Earth's history. Recovery from that event took about 30 million years.
Currently, human activities are driving a rapid decline in biodiversity, often referred to as the Holocene extinction or the sixth mass extinction. It was estimated in 2007 that up to 30% of all species could be extinct by 2050. Habitat destruction—particularly for agriculture—is a primary driver of this decline. Climate change is also a major contributor, affecting entire biomes. This anthropogenic extinction may have begun during the late Pleistocene, as some studies suggest that the extinction that took place around the end of the last ice age partly resulted from overhunting.
Biodiversity as a term is often used in a broader replacement for more specific terms like species diversity and species richness. However, the definition of biodiversity remains fluid and continues to evolve. For instance, the Food and Agriculture Organization of the United Nations (FAO) defined biodiversity in 2019 as "the variability that exists among living organisms (both within and between species) and the ecosystems of which they are part." The World Health Organization updated their website's definition of biodiversity to be the "variability among living organisms from all sources." These definitions reflect current understanding while emphasizing the breadth of the concept.
Since the rate of extinction has increased, many extant species may become extinct before they are described. Not surprisingly, in the the most studied groups are birds and mammals, whereas fishes and arthropods are the least studied animal groups.
In absolute terms, the planet has lost 58% of its biodiversity since 1970 according to a 2016 study by the World Wildlife Fund. The Living Planet Report 2014 claims that "the number of mammals, birds, reptiles, amphibians, and fish across the globe is, on average, about half the size it was 40 years ago". Of that number, 39% accounts for the terrestrial wildlife gone, 39% for the marine wildlife gone and 76% for the freshwater wildlife gone. Biodiversity took the biggest hit in Latin America, plummeting 83 percent. High-income countries showed a 10% increase in biodiversity, which was canceled out by a loss in low-income countries. This is despite the fact that high-income countries use five times the ecological resources of low-income countries, which was explained as a result of a process whereby wealthy nations are outsourcing resource depletion to poorer nations, which are suffering the greatest ecosystem losses.
A 2017 study published in PLOS One found that the biomass of insect life in Germany had declined by three-quarters in the last 25 years. Dave Goulson of Sussex University stated that their study suggested that humans "appear to be making vast tracts of land inhospitable to most forms of life, and are currently on course for ecological Armageddon. If we lose the insects then everything is going to collapse."
In 2020 the World Wildlife Foundation published a report saying that "biodiversity is being destroyed at a rate unprecedented in human history". The report claims that 68% of the population of the examined species were destroyed in the years 1970 – 2016.
Of 70,000 monitored species, around 48% are experiencing population declines from human activity (in 2023), whereas only 3% have increasing populations.
of decline in biodiversity in the current sixth mass extinction match or exceed rates of loss in the five previous Extinction event in the fossil record. Biodiversity loss is in fact "one of the most critical manifestations of the Anthropocene" (since around the 1950s); the continued decline of biodiversity constitutes "an unprecedented threat" to the continued existence of human civilization. The reduction is caused primarily by human impacts, particularly habitat destruction.
Since the Stone Age, species loss has accelerated above the average basal rate, driven by human activity. Estimates of species losses are at a rate 100–10,000 times as fast as is typical in the fossil record.
Loss of biodiversity results in the loss of natural capital that supplies ecosystem goods and services. Species today are being wiped out at a rate 100 to 1,000 times higher than baseline, and the rate of extinctions is increasing. This process destroys the resilience and adaptability of life on Earth.UK Government Official Documents, February 2021, "The Economics of Biodiversity: The Dasgupta Review Headline Messages" p. 1
In 2006, many species were formally classified as rare species or endangered or threatened; moreover, scientists have estimated that millions more species are at risk which have not been formally recognized. About 40 percent of the 40,177 species assessed using the IUCN Red List criteria are now listed as threatened with extinction—a total of 16,119. As of late 2022 9251 species were considered part of the IUCN's critically endangered.
Numerous scientists and the IPBES Global Assessment Report on Biodiversity and Ecosystem Services assert that human population growth and overconsumption are the primary factors in this decline. However, other scientists have criticized this finding and say that loss of habitat caused by "the growth of commodities for export" is the main driver. A 2025 study found that human activities are responsible for biodiversity loss across all species and ecosystems.
Some studies have however pointed out that habitat destruction for the expansion of agriculture and the overexploitation of wildlife are the more significant drivers of contemporary biodiversity loss, not climate change.
Diversity consistently measures higher in the tropics and in other localized regions such as the Cape Floristic Region and lower in polar regions generally. Rain forests that have had wet climates for a long time, such as Yasuní National Park in Ecuador, have particularly high biodiversity.
There is local biodiversity, which directly impacts daily life, affecting the availability of fresh water, food choices, and fuel sources for humans. Regional biodiversity includes habitats and ecosystems that synergizes and either overlaps or differs on a regional scale. National biodiversity within a country determines the ability for a country to thrive according to its habitats and ecosystems on a national scale. Also, within a country, endangered species are initially supported on a national level then internationally. Ecotourism may be utilized to support the economy and encourages tourists to continue to visit and support species and ecosystems they visit, while they enjoy the available amenities provided. International biodiversity impacts global livelihood, food systems, and health. Problematic pollution, over consumption, and climate change can devastate international biodiversity. Nature-based solutions are a critical tool for a global resolution. Many species are in danger of becoming extinct and need world leaders to be proactive with the Kunming-Montreal Global Biodiversity Framework.
Terrestrial biodiversity is thought to be up to 25 times greater than ocean biodiversity. Forests harbour most of Earth's terrestrial biodiversity. The conservation of the world's biodiversity is thus utterly dependent on the way in which we interact with and use the world's forests. (2025). 9789251327074, FAO & UNEP. ISBN 9789251327074 Text was added from this source which has a Wikipedia-specific A new method used in 2011, put the total number of species on Earth at 8.7 million, of which 2.1 million were estimated to live in the ocean. However, this estimate seems to under-represent the diversity of microorganisms. Forests provide habitats for 80 percent of amphibian species, 75 percent of bird species and 68 percent of mammal species. About 60 percent of all vascular plants are found in tropical forests. Mangroves provide breeding grounds and nurseries for numerous species of fish and shellfish and help trap sediments that might otherwise adversely affect seagrass beds and coral reefs, which are habitats for many more marine species. Forests span around 4 billion acres (nearly a third of the Earth's land mass) and are home to approximately 80% of the world's biodiversity. About 1 billion hectares are covered by primary forests. Over 700 million hectares of the world's woods are officially protected.
The biodiversity of forests varies considerably according to factors such as forest type, geography, climate and soils – in addition to human use. Most forest habitats in temperate regions support relatively few animal and plant species and species that tend to have large geographical distributions, while the montane forests of Africa, South America and Southeast Asia and lowland forests of Australia, coastal Brazil, the Caribbean islands, Central America and insular Southeast Asia have many species with small geographical distributions. Areas with dense human populations and intense agricultural land use, such as Europe, parts of Bangladesh, China, India and North America, are less intact in terms of their biodiversity. Northern Africa, southern Australia, coastal Brazil, Madagascar and South Africa, are also identified as areas with striking losses in biodiversity intactness. European forests in EU and non-EU nations comprise more than 30% of Europe's land mass (around 227 million hectares), representing an almost 10% growth since 1990.
Even though terrestrial biodiversity declines from the equator to the poles, some studies claim that this characteristic is unverified in aquatic ecosystems, especially in marine ecosystems. The latitudinal distribution of parasites does not appear to follow this rule. (2010). 9780199561353, Oxford University Press. ISBN 9780199561353 Also, in terrestrial ecosystems the soil bacterial diversity has been shown to be highest in temperate climatic zones, and has been attributed to carbon inputs and habitat connectivity.
In 2016, an alternative hypothesis ("the fractal biodiversity") was proposed to explain the biodiversity latitudinal gradient. In this study, the species pool size and the fractal nature of ecosystems were combined to clarify some general patterns of this gradient. This hypothesis considers temperature, moisture, and net primary production (NPP) as the main variables of an ecosystem niche and as the axis of the ecological hypervolume. In this way, it is possible to build fractal hyper volumes, whose fractal dimension rises to three moving towards the equator.
Brazil's Atlantic Forest is considered one such hotspot, containing roughly 20,000 plant species, 1,350 vertebrates and millions of insects, about half of which occur nowhere else. The island of Madagascar and India are also particularly notable. Colombia is characterized by high biodiversity, with the highest rate of species by area unit worldwide and it has the largest number of endemics (species that are not found naturally anywhere else) of any country. About 10% of the species of the Earth can be found in Colombia, including over 1,900 species of bird, more than in Europe and North America combined, Colombia has 10% of the world's mammals species, 14% of the amphibian species and 18% of the bird species of the world. Madagascar dry deciduous forests and lowland rainforests possess a high ratio of endemism. Since the island separated from mainland Africa 66 million years ago, many species and ecosystems have evolved independently. Indonesia's 17,000 islands cover and contain 10% of the world's , 12% of mammals and 17% of , and —along with nearly 240 million people. Many regions of high biodiversity and/or endemism arise from specialized which require unusual adaptations, for example, alpine climate environments in high , or peat .
Accurately measuring differences in biodiversity can be difficult. Selection bias amongst researchers may contribute to biased empirical research for modern estimates of biodiversity. In 1768, Rev. Gilbert White succinctly observed of his Selborne "all nature is so full, that that district produces the most variety which is the most examined."
Biodiversity grew fast during the Phanerozoic (the last 540 million years), especially during the so-called Cambrian explosion—a period during which nearly every phylum of multicellular organisms first appeared. However, recent studies suggest that this diversification had started earlier, at least in the Ediacaran, and that it continued in the Ordovician. Over the next 400 million years or so, invertebrate diversity showed little overall trend and vertebrate diversity shows an overall exponential trend. This dramatic rise in diversity was marked by periodic, massive losses of diversity classified as mass extinction events. A significant loss occurred in anamniotic limbed vertebrates when rainforests collapsed in the Carboniferous, but seem to have been little affected by this event; their diversification slowed down later, around the Asselian/Sakmarian boundary, in the early Cisuralian (Early Permian), about 293 Ma ago. The worst was the Permian-Triassic extinction event, 251 million years ago. Vertebrates took 30 million years to recover from this event.
The most recent major mass extinction event, the Cretaceous–Paleogene extinction event, occurred 66 million years ago. This period has attracted more attention than others because it resulted in the extinction of the , which were represented by many lineages at the end of the Maastrichtian, just before that extinction event. However, many other taxa were affected by this crisis, which affected even marine taxa, such as ammonites, which also became extinct around that time.
The biodiversity of the past is called Paleobiodiversity. The fossil record suggests that the last few million years featured the greatest biodiversity in history. However, not all scientists support this view, since there is uncertainty as to how strongly the fossil record is biased by the greater availability and preservation of recent geology sections. Some scientists believe that corrected for sampling artifacts, modern biodiversity may not be much different from biodiversity 300 million years ago, whereas others consider the fossil record reasonably reflective of the diversification of life. Estimates of the present global macroscopic species diversity vary from 2 million to 100 million, with a best estimate of somewhere near 9 million, the vast majority . Diversity appears to increase continually in the absence of natural selection.
It also appears that the diversity continues to increase over time, especially after mass extinctions.
On the other hand, changes through the Phanerozoic correlate much better with the hyperbolic model (widely used in population biology, demography and macrosociology, as well as fossil biodiversity) than with exponential and logistic models. The latter models imply that changes in diversity are guided by a first-order positive feedback (more ancestors, more descendants) and/or a negative feedback arising from resource limitation. Hyperbolic model implies a second-order positive feedback. Differences in the strength of the second-order feedback due to different intensities of interspecific competition might explain the faster rediversification of ammonoids in comparison to bivalves after the end-Permian extinction. The hyperbolic pattern of the world population growth arises from a second-order positive feedback between the population size and the rate of technological growth. The hyperbolic character of biodiversity growth can be similarly accounted for by a feedback between diversity and community structure complexity. The similarity between the curves of biodiversity and human population probably comes from the fact that both are derived from the interference of the hyperbolic trend with cyclical and stochastic dynamics.
Most biologists agree however that the period since human emergence is part of a new mass extinction, named the Holocene extinction event, caused primarily by the impact humans are having on the environment. National Survey Reveals Biodiversity Crisis American Museum of Natural History It has been argued that the present rate of extinction is sufficient to eliminate most species on the planet Earth within 100 years.
New species are regularly discovered (on average between 5–10,000 new species each year, most of them ) and many, though discovered, are not yet classified (estimates are that nearly 90% of all are not yet classified). Most of the terrestrial diversity is found in and in general, the land has more species than the ocean; some 8.7 million species may exist on Earth, of which some 2.1 million live in the ocean.
The age of Earth is about 4.54 billion years. The earliest undisputed evidence of life dates at least from 3.7 billion years ago, during the Eoarchean era after a geological crust started to solidify following the earlier molten Hadean eon. (2025). 9780071122610, McGraw-Hill. ISBN 9780071122610 There are microbial mat fossils found in 3.48 billion-year-old sandstone discovered in Western Australia. Other early physical evidence of a biogenic substance is graphite in 3.7 billion-year-old Metasediment discovered in Western Greenland. More recently, in 2015, "remains of Biotic material" were found in 4.1 billion-year-old rocks in Western Australia. According to one of the researchers, "If life arose relatively quickly on Earth...then it could be common in the universe."
Ecosystem services have been grouped in three types:
Experiments with controlled environments have shown that humans cannot easily build ecosystems to support human needs; for example entomophily cannot be mimicked, though there have been attempts to create artificial pollinators using unmanned aerial vehicles. The economic activity of pollination alone represented between $2.1–14.6 billion in 2003. Other sources have reported somewhat conflicting results and in 1997 Robert Costanza and his colleagues reported the estimated global value of ecosystem services (not captured in traditional markets) at an average of $33 trillion annually.
Greater species diversity
Agricultural diversity can also be divided by whether it is 'planned' diversity or 'associated' diversity. This is a functional classification that we impose and not an intrinsic feature of life or diversity. Planned diversity includes the crops which a farmer has encouraged, planted or raised (e.g. crops, covers, symbionts, and livestock, among others), which can be contrasted with the associated diversity that arrives among the crops, uninvited (e.g. herbivores, weed species and pathogens, among others).
Associated biodiversity can be damaging or beneficial. The beneficial associated biodiversity include for instance wild pollinators such as wild bees and syrphid flies that pollinate crops and natural enemies and antagonists to pests and pathogens. Beneficial associated biodiversity occurs abundantly in crop fields and provide multiple ecosystem services such as pest control, nutrient cycling and pollination that support crop production.
Although about 80 percent of humans' food supply comes from just 20 kinds of plants, humans use at least 40,000 species. Earth's surviving biodiversity provides resources for increasing the range of food and other products suitable for human use, although the present extinction rate shrinks that potential.
Some of the health issues influenced by biodiversity include dietary health and nutrition security, infectious disease, medical science and medicinal resources, social and psychological health. Biodiversity is also known to have an important role in reducing disaster risk, including rising sea levels. For example, wetland ecosystems along coastal communities serve as excellent water filtration systems, storage, and ultimately create a buffer region between the ocean and mainland neighborhoods in order to prevent water reaching these communities under climate change pressures or storm storages. Other examples of diverse species or organisms are present around the world, offering their resourceful utilities to provide protection of human survival.
Biodiversity provides critical support for drug discovery and the availability of medicinal resources. (2006) "Molecular Pharming" GMO Compass Retrieved 5 November 2009, GMOcompass.org A significant proportion of drugs are derived, directly or indirectly, from biological sources: at least 50% of the pharmaceutical compounds on the US market are derived from plants, animals and , while about 80% of the world population depends on medicines from nature (used in either modern or traditional medical practice) for primary healthcare. Only a tiny fraction of wild species has been investigated for medical potential.
are particularly important, especially their chemical and physical properties that have paved the way for numerous pharmaceutical achievements; the immense diversity of marine organisms have led to scientific discoveries including medical treatments to cancer, viral bacteria, AIDS, etc. This process of bioprospecting can increase biodiversity loss, as well as violating the laws of the communities and states from which the resources are taken.
Biodiversity's economic resources are worth at around $150 trillion annually which is roughly twice the world's GDP. The loss of biodiversity is actually harming the GDP of the world by costing an estimated $5 trillion annually.
Business supply chains rely heavily on ecosystems remaining relatively maintained and nurtured. A disruption to these supply chains would negatively impact many businesses that would end up costing them more than what they are gaining.
Biodiversity also affords many non-material benefits including spiritual and aesthetic values, knowledge systems and education.
Species are increasingly being moved by humans (on purpose and accidentally). Some studies say that diverse ecosystems are more resilient and resist invasive plants and animals. Many studies cite effects of invasive species on natives, but not extinctions.
Invasive species seem to increase local (alpha diversity) diversity, which decreases turnover of diversity (beta diversity). Overall gamma diversity may be lowered because species are going extinct because of other causes, but even some of the most insidious invaders (e.g.: Dutch elm disease, emerald ash borer, chestnut blight in North America) have not caused their host species to become extinct. Local extinction, population decline and homogenization of regional biodiversity are much more common. Human activities have frequently been the cause of invasive species circumventing their barriers, by introducing them for food and other purposes. Human activities therefore allow species to migrate to new areas (and thus become invasive) occurred on time scales much shorter than historically have been required for a species to extend its range.
At present, several countries have already imported so many exotic species, particularly agricultural and ornamental plants, that their indigenous fauna/flora may be outnumbered. For example, the introduction of kudzu from Southeast Asia to Canada and the United States has threatened biodiversity in certain areas. Another example are , which have invaded forests, shrublands and grasslands in the southern hemisphere.
Conservation biology matured in the mid-20th century as ecologists, naturalists and other scientists began to research and address issues pertaining to global biodiversity declines.
The conservation ethic advocates management of for the purpose of sustaining biodiversity in species, ecosystems, the Evolution and human culture and society.
Conservation biology is reforming around strategic plans to protect biodiversity. Preserving global biodiversity is a priority in strategic conservation plans that are designed to engage public policy and concerns affecting local, regional and global scales of communities, ecosystems and cultures.Example: Gascon, C., Collins, J. P., Moore, R. D., Church, D. R., McKay, J. E. and Mendelson, J. R. III (eds) (2007). Amphibian Conservation Action Plan. IUCN/SSC Amphibian Specialist Group. Gland, Switzerland and Cambridge, UK. 64pp. Amphibians.org , see also Millenniumassessment.org, Europa.eu Action plans identify ways of sustaining human well-being, employing natural capital, macroeconomic policies including economic incentives, and ecosystem services.
In the EU Directive 1999/22/EC zoos are described as having a role in the preservation of the biodiversity of wildlife animals by conducting research or participation in .
As sustainable populations of the remaining native species in an area become assured, "missing" species that are candidates for reintroduction can be identified using databases such as the Encyclopedia of Life and the Global Biodiversity Information Facility.
Protected areas safeguard nature and cultural resources and contribute to livelihoods, particularly at local level. There are over 238 563 designated protected areas worldwide, equivalent to 14.9 percent of the earth's land surface, varying in their extension, level of protection, and type of management (IUCN, 2018).
The benefits of protected areas extend beyond their immediate environment and time. In addition to conserving nature, protected areas are crucial for securing the long-term delivery of ecosystem services. They provide numerous benefits including the conservation of genetic resources for food and agriculture, the provision of medicine and health benefits, the provision of water, recreation and tourism, and for acting as a buffer against disaster. Increasingly, there is acknowledgement of the wider socioeconomic values of these natural ecosystems and of the ecosystem services they can provide.
Both of these serve as places in which biodiversity can be preserved rather than harmed. According to an article published in the National Park Service website, national parks aim their resources at maintaining animal and habitat integrity through conservation and preservation of their ecosystems. This along with educating the general public on wildlife functions, the aim for an increase in biodiversity is one of many goals trying to be focused on through national parks.
There is an estimated 726 million ha of forest in protected areas worldwide. Of the six major world regions, South America has the highest share of forests in protected areas, 31 percent. (2025). 9789251325810, FAO. ISBN 9789251325810 Text was added from this source which has a Wikipedia-specific The play a vital role in harboring more than 45,000 floral and 81,000 faunal species of which 5150 floral and 1837 faunal species are Endemism. In addition, there are 60,065 different tree species in the world. Plant and animal species confined to a specific geographical area are called endemic species.
In forest reserves, rights to activities like hunting and grazing are sometimes given to communities living on the fringes of the forest, who sustain their livelihood partially or wholly from forest resources or products.
Approximately 50 million hectares (or 24%) of European forest land is protected for biodiversity and landscape protection. Forests allocated for soil, water, and other ecosystem services encompass around 72 million hectares (32% of European forest area).
The concept of nature-positive is playing a role in mainstreaming the goals of the Global Biodiversity Framework (GBF) for biodiversity. The aim of mainstreaming is to embed biodiversity considerations into public and private practice to conserve and Sustainability biodiversity on global and local levels. The concept of nature-positive refers to the societal goal to halt and reverse biodiversity loss, measured from a baseline of 2020 levels, and to achieve full so-called "nature recovery" by 2050.
Volunteer observers have made significant contributions to on-the-ground knowledge about biodiversity, and recent improvements in technology have helped increase the flow and quality of occurrences from citizen sources. A 2016 study published in Biological Conservation registers the massive contributions that citizen scientists already make to data mediated by the Global Biodiversity Information Facility (GBIF). Despite some limitations of the dataset-level analysis, it is clear that nearly half of all occurrence records shared through the GBIF network come from datasets with significant volunteer contributions. Recording and sharing observations are enabled by several global-scale platforms, including iNaturalist and eBird. (2025). 9783319272887, Springer Nature. ISBN 9783319272887
Global agreements such as the Convention on Biological Diversity, give "sovereign national rights over biological resources" (not property). The agreements commit countries to "conserve biodiversity", "develop resources for sustainability" and "share the benefits" resulting from their use. Biodiverse countries that allow bioprospecting or collection of natural products, expect a share of the benefits rather than allowing the individual or institution that discovers/exploits the resource to capture them privately. Bioprospecting can become a type of biopiracy when such principles are not respected.
Sovereignty principles can rely upon what is better known as Access and Benefit Sharing Agreements (ABAs). The Convention on Biodiversity implies informed consent between the source country and the collector, to establish which resource will be used and for what and to settle on a fair agreement on benefit sharing.
On the 19 of December 2022, during the 2022 United Nations Biodiversity Conference every country on earth, with the exception of the United States and the Holy See, signed onto the agreement which includes protecting 30% of land and oceans by 2030 (30 by 30) and 22 other targets intended to reduce biodiversity loss. The agreement includes also recovering 30% of earth degraded ecosystems and increasing funding for biodiversity issues.
The EU Biodiversity Strategy for 2030 include the next targets:
Approximately half of the global GDP depend on nature. In Europe many parts of the economy that generate trillions of euros per year depend on nature. The benefits of Natura 2000 alone in Europe are €200 – €300 billion per year.
Uniform approval for use of biodiversity as a legal standard has not been achieved, however. Bosselman argues that biodiversity should not be used as a legal standard, claiming that the remaining areas of scientific uncertainty cause unacceptable administrative waste and increase litigation without promoting preservation goals.
India passed the Biological Diversity Act in 2002 for the conservation of biological diversity in India. The Act also provides mechanisms for equitable sharing of benefits from the use of traditional biological resources and knowledge.
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