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An aril (), also called arillus (plural arilli), is a specialized outgrowth from a that partly or completely covers the seed. An arillode, or false aril, is sometimes distinguished: whereas an aril grows from the attachment point of the seed to the ovary (from the or hilum), an arillode forms from a different point on the seed coat. The term "aril" is sometimes applied to any fleshy appendage of the seed in , such as the mace of the seed. Arils and arillodes are often edible enticements that encourage animals to transport the seed, thereby assisting in seed dispersal.

(1992). 9780521373500, Cambridge University Press. .
Pseudarils are aril-like structures commonly found on the of species that develop from the of the ovary. The fleshy, edible pericarp splits neatly in two halves, then falling away or being eaten to reveal a brightly coloured pseudaril around the black seed.

The aril may create a -like structure, called (among other names) a . False fruit are found in numerous taxa. The edible false fruit of the , and fruits are highly developed arils surrounding the seed rather than a layer. Such arils are also found in a few species of , notably the and related such as the lleuque and the kahikatea. Instead of the typical of most gymnosperms, the reproductive structure of the yew consists of a single seed that becomes surrounded by a fleshy, cup-like covering. This covering is derived from a highly modified cone scale.


Development in Taxus
In European yew plants ( ), the aril starts out as a small, green band at the base of the seed, then turns brown to red as it enlarges and surrounds the seed, eventually becoming fleshy and scarlet in color at maturity. The aril is attractive to fruit-eating and is non-toxic. All other parts of the yew are toxic, including the seed housed inside the aril. If the seed is crushed, breaks or splits in the stomach of a human, bird or another animal, it will result in poisoning. Birds digest the fleshy aril as a food source, and pass the seeds out in their droppings, promoting of the seeds.


In Dacrycarpus dacrydioides
The kahikatea tree, Dacrycarpus dacrydioides, is native to . In pre-European times the aril of the kahikatea was a food source for Māori. The washed arils were called koroi and were eaten raw.


See also
  • , fleshy structures attached to the seeds of many plant species
  • , a fleshy cone borne chiefly by junipers and cypresses
  • , a fleshy epidermal layer of a seed coat, as in pomegranate


Further reading
  • Anderson, E. & Owens, J. N. (2003). Analysing the reproductive biology of Taxus: should it be included in Coniferales? Acta Hort. 615: 233–234.

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