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   » » Wiki: Zapsalis
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Zapsalis is a of . It is a , often considered because of the fragmentary nature of the , which include teeth but no other remains.


Etymology
The generic name is derived from Greek za~, "thorough", and psalis, "pair of scissors". The specific name means "abrading" in .


History and classification
Fossils of Zapsalis were first described by American paleontologist Edward Drinker Cope in 1876 but as species of the large carnivorous theropod Laelaps (now ).Cope, E. D. (1876). Descriptions of some vertebrate remains from the Fort Union beds of Montana. Proceedings of the Academy of natural Sciences of Philadelphia, 248-261. Cope erected 2 species, Laelaps explanatus and L. laevifrons, the former based on a collection of 27 teeth and the latter based on a single tooth. It wasn't until later in 1876 that Cope made the genus Zapsalis, with Z. abradens as the type, based on a second .Cope, E. D. (1876). On some extinct reptiles and Batrachia from the Judith River and Fox Hills beds of Montana. Proceedings of the Academy of natural Sciences of Philadelphia, 340-359. All of the fossils were collected from the age strata of the Judith River Formation in , USA. Cope named Zapsalis during the , his competition with Yale paleontologist Othniel Charles Marsh, to collect and describe as many fossil taxa as possible.Brinkman, P. D. (2010). The second Jurassic dinosaur rush. University of Chicago Press.

After the Bone Wars, the type fossils of Zapsalis and the Laelaps species were sold to the American Museum of Natural History in New York. In the wake of the Bone Wars, the complicated errors in dinosaur taxonomy were left to other paleontologists, with the Laelaps species being moved to other theropod dinosaurs like ,Osborn, H. F., & Lambe, L. M. (1902). On Vertebrata of the Mid-Cretaceous of the North West Territory (Vol. 3). Government Printing Bureau.Lambe, L. M. (1902). New genera and species from the Belly River series (Mid-Cretaceous). Contributions to Canadian Paleontology, Geological Survey of Canada 3: 25-81. 1918. The Cretaceous genus Stegoceras typifying a new family referred provisionally to the Stegosauria. Transactions of the Royal Society of Canada, 12, 23-36. ,Hatcher, J. B. (1903). Osteology of Haplocanthosaurus, with description of a new species and remarks on the probable habits of the Sauropoda and the age and origin of the Atlantosaurus beds: additional remarks on Diplodocus (Vol. 2, No. 1). Carnegie Museum. and .Matthew, W. D., & Brown, B. (1922). The family Deinodontidae, with notice of a new genus from the Cretaceous of Alberta. Bulletin of the AMNH; v. 46, article 6.Kuhn, O. (1939). Saurischia, Fossilium Catalogus I: Animalia, Pars 87. W. Quenstedt, Munich. Z. abradens was moved to Dromaeosaurus and synonymized with the other Dromaeosaur .Hotton III, N. (1965). Fossil Vertebrates from the Late Cretaceous Lance Formation, Eastern Wyoming. It wasn't until 2002 that e.a. concluded the teeth represented a separate " ?Dromaeosaurus Morphotype A". In 2013 and Philip Currie recognised Zapsalis as a valid taxon from the Judith River and Dinosaur Park Formation. The teeth are typified by a combination of rounded denticles, straight rear edge and vertical grooves. Similar teeth from the older Milk River Formation were referred to a cf. Zapsalis. In 2019, Currie and Evans announced that the Zapsalis teeth from the Dinosaur Park Formation represented the second premaxillary tooth of Saurornitholestes langstoni, in a paper describing a complete skull of that species. The authors kept the species distinct because the type species' holotype is likely indeterminate on a species level.Baszio, S. (1997). Systematic palaeontology of isolated dinosaur teeth from the latest Cretaceous of south Alberta, Canada. Courier Forschungsinstitut Senckenberg, 196, 33-77.

As for Laelaps explanatus and L. laevifrons, they were never synonymized with Zapsalis but have been synonymized with Saurornitholestes langstoni and in turn, Zapsalis, as well.


Description
The type tooth of Z. abradens is flat lingually, with no mesial serrations and 3 distal serrations per millimeter and is 12 mm in total length. There are three lingual ridges and four labial ones. Currie & Evans, 2019 diagnosed Zapsalis from Saurornitholestes by noting the type of the former is lacking mesial serrations and being concave apicodistally, and therefore "recommended that the two genera be kept separate." The second teeth of Zapsalis and other dromaeosaurids may have been structurally specialized for preening feathers, as seen in some as well.


Paleoenvironment
All four named species are known from the Judith River Formation, the site of expeditions first by Edward Drinker Cope's crews during the early stages of the , including the discoveries of many taxa that he named, though all are now seen as dubious. These include fossils of large, carnivorous theropods like and . As for the herbivorous , like the beaked and were named. The most common fossils are those of the horned like , , and . Lastly, the armored is known from scattered teeth.Cope, E.D. (1879). Hayden, F.V. (ed.). "The Relations of the Horizons of Extinct Vertebrata". United States Geological and Geographical Survey. 5 (1): 37–38.


See also
  • Timeline of dromaeosaurid research

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