Pyrococcus is a genus of .[See the NCBI webpage on Pyrococcus. Data extracted from the ]
Description and significance
Pyrococcus has similar characteristics of other thermoautotrophican
archaea such as
Archaeoglobus, and
Methanococcus in the respect that they are all
thermophile and anaerobic.
Pyrococcus differs, however, because its optimal growth temperature is nearly 100 °C and dwells at a greater sea depth than the other archaea. Studying
Pyrococcus helps give insight to possible mechanisms used to endure extreme environmental conditions like high temperatures and high pressure.
Phylogeny
Genome structure
Three of the
Pyrococcus species have been sequenced.
P. furiosus is the largest containing 1.9 Mb followed by
P. abyssi with 1.8 Mb and
P. horikoshii with 1.7 Mb. The
encode for many different metabolic
which gives themselves a wider spectrum of living conditions because they can transport and metabolize a wide range of organic substances. Variation was detected between species as well.
Cell structure and metabolism
The cells of
Pyrococcus are about 0.8–2 μm and are slightly irregular cocci in shape. They show a polar grouping of
Archaellum and are enveloped by an
S-layer enclosing a
space around the cytoplasmic membrane.
Pyrococcus species are anaerobic but vary slightly concerning their metabolism.
Peptide fermentation is the principle metabolic pathway; however, growth has been observed for
P. furiosus and
P. abyssi on
starch,
maltose, and
pyruvate, but not for
P. horikoshii. While the presence of elemental sulfur is not needed for growth, growth is enhanced with the addition of S
o.
Ecology
Pyrococcus species inhabit environments with extremely high temperatures such as hydrothermal vents. Optimal growth conditions include a
pH level of about 7, a salt concentration around 2.5%, and a temperature around 98 °C. Growing in temperatures this high, it is easy to see why they are anaerobic since at these boiling temperatures hardly any oxygen will be available. In the example of Hydrothermal vents, where
P. abyssi has been found, there is no sunlight and the pressure is around 200 atm in addition to the extremely high temperature.
See also
Further reading
