Nemastomatidae

 ( Harvestman Families ) 

For a broader recent listing of species see:

• Acromitostoma Roewer, 1951

* Acromitostoma hispanum (Roewer, 1917)Spain

* Acromitostoma rhinoceros Roewer, 1951Spain, Morocco

• Burnia Prieto, 2021

* Burnia sexmucronata (Simon, 1911)Spain

* Burnia spelaea Prieto, 2021Spain

• Carinostoma Kratochvíl, 1958

* Carinostoma carinatum (Roewer, 1914)southeastern Europe

* Carinostoma elegans (Sørensen, 1894)southeastern Europe

* Carinostoma ornatum (Hadzi, 1940)southeastern Europe

• Caucnemastoma Martens, 2006

* Caucnemastoma golovatchi Martens, 2006Russia

* Caucnemastoma martensi Snegovaya, 2011Russia

• Centetostoma Kratochvíl, 1958

* Centetostoma centetes (Simon, 1881)France (Alps)

* Centetostoma juberthiei Martens, 2011France/Spain (Pyrenees)

* Centetostoma scabriculum (Simon, 1879)France/Spain (Pyrenees)

* Centetostoma ventalloi (Mello-Leitao, 1936)France/Spain (Pyrenees)

• Giljarovia Kratochvíl, 1958

* Giljarovia crimeana Tchemeris & Kovblyuk, 2012Ukraine

* Giljarovia kratochvili Snegovaya, 2011Russia

* Giljarovia redikorzevi Charitonov, 1946Georgia

* Giljarovia rossica Kratochvíl, 1958Russia

* Giljarovia stridula (Kratochvíl, 1958)Russia

* Giljarovia tenebricosa (Redikortsev, 1936)Georgia, Azerbaijan, Turkey

* Giljarovia thoracocornuta Martens, 2006Russia

* Giljarovia triangula Martens, 2006Russia, Georgia

* Giljarovia trianguloides Martens, 2006Russia

* Giljarovia turcica Gruber, 1976Turkey

* Giljarovia vestita Martens, 2006Russia, Georgia

• Hadzinia Šilhavý, 1966

* Hadzinia ferrani Novak & Kozel, 2014Slovenia

* Hadzinia karamani (Hadzi, 1940)Bosnia & Herzegovina, Croatia

• Histricostoma Kratochvíl, 1958

* Histricostoma anatolicum (Roewer, 1962)Turkey

* Histricostoma argenteolunulatum (Canestrini, 1875)France (Corsica), Italy, Greece

* Histricostoma caucasicum (Redikorzev, 1936)Russia

* Histricostoma creticum (Roewer, 1928)Greece

* Histricostoma dentipalpe (Anton Ausserer, 1867)Central Europe

* Histricostoma drenskii Kratochvíl, 1958Bulgaria

* Histricostoma gruberi Snegovaya & Marusik, 2012Turkey

* Histricostoma mitovi Snegovaya & Marusik, 2012Turkey

• Mediostoma Kratochvíl, 1958

* Mediostoma armatum Martens, 2006Iran

* Mediostoma ceratocephalum Gruber, 1976Turkey

* Mediostoma cypricum (Roewer, 1951)Greece/Turkey: Cyprus, Israel, Lebanon,

* Mediostoma globuliferum (L. Koch, 1867)Greece

* Mediostoma humerale (C.L. Koch, 1839)Greece, Albania

* Mediostoma izmirica Snegovaya, Kurt, & Yağmur, 2016Turkey

* Mediostoma nigrum Martens, 2006Iran, Azerbaijan

* Mediostoma pamiricum Staręga, 1986Tajikistan

* Mediostoma stussineri (Simon, 1885) Greece

* Mediostoma talischense (Morin, 1937)Azerbaijan

* Mediostoma variabile Martens, 2006Iran, Azerbaijan

* Mediostoma vityae (Roewer, 1927)Greece

• Mitostoma Roewer, 1951

* Mitostoma alpinum (Hadži, 1931)Southeastern Europe

* Mitostoma anophthalmum (Fage, 1946)Italy

* Mitostoma atticum (Roewer, 1928)Greece

* Mitostoma cancellatum (Roewer, 1917)Balkans

* Mitostoma carneluttii Hadži, 1973Montenegro

* Mitostoma chrysomelas (Hermann, 1804)Europe (widespread)

* Mitostoma daccordii Tedeschi & Sciaky, 1997Italy

* Mitostoma fabianae Tedescie & Sciaky, 1997Italy

* Mitostoma gracile (Redikortsev, 1936)Russia, Georgia, etc.

* Mitostoma macedonicum Hadži, 1973Macedonia

* Mitostoma olgae (Šilhavý, 1939)(see below Balkans)

:* Mitostoma olgae decorum (Šilhavý, 1939)Montenegro

:* Mitostoma olgae kratochvili (Šilhavý, 1939)Croatia

:* Mitostoma olgae olgae (Šilhavý, 1939)Montenegro

:* Mitostoma olgae zorae Hadži, 1973Macedonia

* Mitostoma orobicum (Caporiacco, 1949)Italy

* Mitostoma patrizii Roewer, 1953Italy

* Mitostoma pyrenaeum (Simon, 1879)France/Spain (Pyrenees)

* Mitostoma sabbadinii Tedeschi & SciakyItaly

* Mitostoma valdemonense Marcellino, 1974Italy

* Mitostoma zmajevicae Hadži, 1973Macedonia

• Nemaspela Silhavý, 1966

* Nemaspela abchasica Ljovuschkin & Starobogatov, 1963Russia, Georgia/Abkhazia

* Nemaspela birsteini Levushkii, 1972Georgia/Abkhazia

* Nemaspela borkoae Kozel, Delić & Novak, 2020Montenegro

* Nemaspela caeca (Grese, 1911)Ukraine (Crimea) Can.

* Nemaspela femorecurvata Martens, 2006Georgia

* Nemaspela gagrica Tchemeris, 2013Georgia/Abkhazia

* Nemaspela kovali Tchemeris, 2009Russia

* Nemaspela ladae Karaman, 2013Bosnia & Herzegovina

* Nemaspela melouri Martens, Maghradze & Barjadze, 2021Georgia

* Nemaspela prometheusi Martens, Maghradze & Barjadze, 2021Georgia

* Nemaspela sokolovi (Ljovuschkin & Starobogatov, 1963)Russia

* Nemaspela taurica (Lebedinski, 1914)Ukraine (Crimea) (See as synonym under Nemaspela caeca per some e.g. Chemeris (2009).

• Nemastoma C. L. Koch, 1836

* Nemastoma bidentatum Roewer, 1914Europe

:* Nemastoma bidentatum bidentatum Roewer, 1914Central Europe, Southeastern Europe

:* Nemastoma bidentatum gruberi Novak, Slana Novak, Kozel & Raspotnig, 2021Southern Europe

:* Nemastoma bidentatum martensi Novak, Slana Novak & Raspotnig, 2021Slovenia

:* Nemastoma bidentatum schmidti Novak, Raspotnig & Slana Novak, 2021Central Europe, Southern Europe

:* Nemastoma bidentatum sneznikensis Novak, Komposch, Slana Novak & Raspotnig, 2021Slovenia

:* Nemastoma bidentatum sparsum Gruber & Martens, 1968Central Europe, Southeastern Europe

* Nemastoma bimaculatum (Fabricius, 1775)Europe (Widespread), Iceland

* Nemastoma daciscum Koch, 1869(?) ''Insertae.

* Nemastoma dentigerum Canestrini, 1873Europe (Widespread)

* Nemastoma kozari Novak, Kozel, Podlesnik & Raspotnig, 2021Bosnia & Herzegovina

* Nemastoma lilliputanum (Lucas, 1846)Algeria ''Insertae.

* Nemastoma lugubre (Müller, 1776)Europe (Widespread)

* Nemastoma pluridentatum (Hadži, 1973)Balkans

* Nemastoma relictum (Gruber & Martens, 1968)Austria

* Nemastoma rude Simon, 1881France ''Insertae.

* Nemastoma schuelleri Gruber & Martens, 1968Austria

* Nemastoma transsylvanicum Gruber & Martens, 1968Romania

* Nemastoma triste (C.L. Koch, 1835)Central Europe

• Nemastomella Mello-Leitão, 1936

* Nemastomella armatissima (Roewer, 1962)Portugal

* Nemastomella bacillifera (Simon, 1879)Western Europe

:* Nemastomella bacillifera bacillifera (Simon, 1879)France/Spain (Pyrenees)

:* Nemastomella bacillifera carbonaria (Simon, 1907)Spain

* Nemastomella cristinae (Rambla, 1969)Spain

* Nemastomella dentipatellae (Dresco, 1967)Spain

* Nemastomella dipentata (Rambla, 1959)Spain

* Nemastomella dubia (Mello-Leitão, 1936)Spain Inc..

* Nemastomella gevia Prieto, 2004Spain

* Nemastomella hankiewiczii (Kulczyński, 1909)Portugal

* Nemastomella iberica (Rambla in Dresco, 1967)Spain

* Nemastomella maarebensis (Simon, 1913)Algeria

* Nemastomella manicata (Simon, 1913)Spain

* Nemastomella monchiquensis (Kraus, 1961)Portugal

* Nemastomella spinosissima (Kraus, 1961)Spain

• Paranemastoma Redikorzev, 1936

* Paranemastoma aeginum (Roewer, 1951)Greece ''Nomen

* Paranemastoma amseli (Roewer, 1951)Italy ''Nomen

* Paranemastoma amuelleri (Roewer, 1951)Macedonia ''Nomen

* Paranemastoma ancae Avram, 1973Romania

* Paranemastoma armatum (Kulczyński, 1909)Balkans

* Paranemastoma aurigerum (Roewer, 1951)Bulgaria (all subspecies)

:* Paranemastoma aurigerum aurigerum (Roewer, 1951)Bulgaria

:* Paranemastoma aurigerum joannae Staręga, 1976Bulgaria

:* Paranemastoma aurigerum ryla (Roewer, 1951)Bulgaria

* Paranemastoma aurosum (L. Koch, 1869)Greece

* Paranemastoma bacurianum (Mkheidze, 1959)Georgia

* Paranemastoma beroni Mitov, 2011Bulgaria

* Paranemastoma bicuspidatum (C.L. Koch, 1835)Central Europe

* Paranemastoma bolei (Hadži, 1973)Macedonia ''Nomen

* Paranemastoma brevipalpatum (Roewer, 1951)Spain ''Nomen

* Paranemastoma bureschi (Roewer, 1926)Bulgaria, Tunisia (? )

* Paranemastoma caporiaccoi (Roewer, 1951)Italy ''Nomen

* Paranemastoma carneluttii (Hadži, 1973)Montenegro ''Nomen

* Paranemastoma corcyraeum (Roewer, 1917)Balkans

* Paranemastoma emigratum (Roewer, 1959India (?) ''Nomen

* Paranemastoma ferkeri (Roewer, 1951)Turkey ''Nomen

* Paranemastoma filipes (Roewer, 1917)South Caucasus, Iran

* Paranemastoma gostivarense (Hadži, 1973)Macedonia ''Nomen

* Paranemastoma ikarium (Roewer, 1951)Greece ''Nomen

* Paranemastoma ios (Roewer, 1917)Greece ''Nomen

* Paranemastoma iranicum Martens, 2006Iran

* Paranemastoma kaestneri (Roewer, 1951)Greece ''Nomen

* Paranemastoma kalischevskyi (Roewer, 1951)South Caucasus

* Paranemastoma karolianum Çorak Öcal, Bayram, Yiğit & Sancak, 2017Turkey

* Paranemastoma kochii (Nowicki, 1870)Central Europe, Southeastern Europe

* Paranemastoma longipalpatum (Roewer, 1951)Greece ''Nomen

* Paranemastoma longipes (Schenkel, 1947)Southeastern Europe

* Paranemastoma macedonicum (Hadži, 1973)Macedonia ''Nomen

* Paranemastoma machadoi (Roewer, 1951)Spain ''Nomen

* Paranemastoma mackenseni (Roewer, 1923)Serbia ''Nomen

* Paranemastoma montenigrinum (Nosek, 1904)Montenegro ''Nomen

* Paranemastoma monticola (Babalean, 2011)Romania

* Paranemastoma multisignatum (Hadži, 1973)Macedonia ''Nomen

* Paranemastoma nigrum (Hadži, 1973)Macedonia ''Nomen

* Paranemastoma perfugium (Roewer, 1951)Italy ''Nomen

* Paranemastoma quadripunctatum (Perty, 1833)Europe (widespread)

* Paranemastoma radewi (Roewer, 1926)Southeastern Europe

* Paranemastoma redikorzevi (Roewer, 1951)Ukraine ''Nomen

* Paranemastoma santorinum (Roewer, 1951)Greece ''Nomen

* Paranemastoma senussium (Roewer, 1951)Lybia ''Nomen

* Paranemastoma sillii (Herman, 1871)Central Europe, Southeastern Europe

* Paranemastoma simplex (Giltay, 1932)Greece

* Paranemastoma sketi (Hadži, 1973)Montenegro ''NomenCentral Europe, Southeastern Europe

* Paranemastoma spinosulum (Koch, 1869)Greece ''Nomen

* Paranemastoma superbum Redikorzev, 1936Georgia, Turkey

* Paranemastoma thessalum (Simon, 1885)Greece, Bosnia & Herzegovina (?)

* Paranemastoma titaniacum (Roewer, 1914)Southeastern Europe

* Paranemastoma umbo (Roewer, 1951)Georgia ''Nomen

* Paranemastoma werneri (Kulczyński, 1903)Turkey

• Pyza Starega, 1976

* Pyza anatolica (Roewer, 1959)Turkey

* Pyza bosnica (Roewer, 1919)Balkans

* Pyza navarrense (Roewer, 1951)Greece

* Pyza taurica Gruber, 1979Turkey

• Saccarella Schönhofer & Martens, 2012

* Saccarella schilleri Schönhofer & Martens, 2012Italy

• Sinostoma Martens, 2016

* Sinostoma yunnanicum Martens, 2016China

• Starengovia Snegovaya, 2010

* Starengovia ivanloebli Martens, 2017Pakistan

* Starengovia kirgizica Snegovaya, 2010Kyrgyzstan

* Starengovia quadrituberculata Zhang & Martens, 2018China

• Vestiferum Martens, 2006

* Vestiferum alatum Martens, 2006Georgia, Turkey

* Vestiferum funebre (Redikorzev, 1936)Georgia/Abkhazia, Russia (also possibly turkey)

• Asiolasma Martens, 2009 (6 species, Eastern Asia)

* Asiolasma ailaoshan (Zhang, Zhao & Zhang, 2018) China

* Asiolasma angka (Schwendinger & Gruber, 1992)Thailand

* Asiolasma billsheari Martens, 2019China

* Asiolasma damingshan (Zhang & Zhang, 2013)China

* Asiolasma juergengruberi Martens, 2019China

* Asiolasma schwendingeri Martens, 2019Vietnam

• Cladolasma Suzuki, 1963 (1 species, Japan)

* Cladolasma parvulum Suzuki, 1963Japan

• Cryptolasma Cruz-López, Cruz-Bonilla & Francke, 2018 (2 species, Eastern Mexico)

* Cryptolasma aberrante Cruz-López, Cruz-Bonilla & Francke, 2018Mexico

* Cryptolasma citlaltepetl Cruz-López, Cruz-Bonilla & Francke, 2018Mexico

• Dendrolasma Banks, 1894 (1 species, Western USA)

* Dendrolasma dentipalpe Shear & Gruber, 1983USA (California)

* Dendrolasma mirabile Banks, 1894USA (Washington, etc), Canada

• Martensolasma Shear, 20064 (2 species, (Central Mexico)

* Martensolasma catrina Cruz-López, 2017Mexico

* Martensolasma jocheni Shear, 2006Mexico

• Ortholasma Banks, 1894 (5 species, Western USA, Northwestern Mexico)

* Ortholasma colossus Shear, 2010USA (California)

* Ortholasma coronadense Cockerell, 1916USA (California), Mexico (Coronados Islands)

* Ortholasma levipes Shear & Gruber, 1983USA (California)

* Ortholasma pictipes Banks, 1911USA (California)

* Ortholasma rugosum Banks, 1894USA (California)

• Trilasma Goodnight & Goodnight, 1942 (9 species, Mexico, Honduras)

* Trilasma bolivari Goodnight & Goodnight, 1942Mexico

* Trilasma chipinquense Shear, 2010Mexico

* Trilasma hidalgo Shear, 2010Mexico

* Trilasma petersprousei Shear, 2010Mexico

* Trilasma ranchonuevo Shear, 2010Mexico

* Trilasma sbordonii (Šilhavý, 1973)Mexico

* Trilasma tempestado Shear, 2010Mexico

* Trilasma trispinosum Shear, 2010Mexico

* Trilasma tropicum Shear, 2010Honduras

:* † Nemastoma tuberculatum Koch & Berendt, 1854 — fossil: Oligocene

:* † Nemastoma succineum Roewer, 1939 — fossil: Baltic amber

* † Rhabdotarachnoides Haupt, 1956 — fossil

:* † Rhabdotarachnoides simoni Haupt, 1956 — Permian

• (eds.) (2007): Harvestmen - The Biology of Opiliones. Harvard University Press

• (eds.) (2007). Harvestmen - The Biology of Opiliones. Harvard University Press, USA.

For older species listings, see:

• Joel Hallan's Biology Catalog (2005) defunct: target="_blank" rel="nofollow"> Nemastomatidae