Leptostraca (from the Greek language words for thin and shell) is an order of small, marine . Its members, including the well-studied Nebalia, occur throughout the world's oceans and are usually considered to be filter feeder. It is the only extant order in the subclass Phyllocarida. They are believed to represent the most primitive members of their class, the Malacostraca, and first appear in the fossil record during the Cambrian period.[
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Description
Leptostracans are usually small, typically long, but the largest species ( Nebaliopsis) can reach 4 cm, and the Silurian Ceratiocaris could grow to 75 cm.[ Invertebrate Palaeontology and Evolution] They are distinguished from all other members of their class in having seven abdomen segments, instead of six. Their head has stalked , two pairs of antennae (one biramous, one uniramous), and a pair of mandibles but no . They are the only with a carapace that comprises two valves. It covers the head and the thorax, including most of the thoracic appendages, and serves as a brood pouch for the developing . Its anterior tip bears a movable rostrum. Also unique among malacostracans is their eight pairs of thoracic appendages which have been specialized into leaf-like filter feeding organs, and are not used for locomotion. The first six abdominal segments bear , while the seventh bears a pair of caudal furcae, which may be homologous to of other crustaceans.[ A phylogeny of the Leptostraca (Crustacea) with keys to families and genera][ Multicellular Animals: Volume II: The Phylogenetic System of the Metazoa]
Leptostracans have gills on their thoracic limbs, but also breathe through a respiratory membrane on the inside of the carapace. The eggs hatch as a , or "manca" stage, which lacks a fully developed carapace, but otherwise resembles the adult.
Classification
It is now accepted that leptostracans belong to the Malacostraca, and the sister crown group to Leptostraca is Eumalacostraca.
The order Leptostraca is divided into three families, with ten genus containing a total of around 40 validly described extant species:
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