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The hippopotamus ( Hippopotamus amphibius; ; : hippopotamuses), often shortened to hippo (: hippos), further qualified as the common hippopotamus, Nile hippopotamus and river hippopotamus, is a large semiaquatic Mammal native to sub-Saharan Africa. It is one of only two extant taxon species in the family Hippopotamidae, the other being the pygmy hippopotamus ( Choeropsis liberiensis or Hexaprotodon liberiensis). Its name comes from the ancient Greek for "river horse" (ἱπποπόταμος).
After and , the hippopotamus is the next largest land mammal. It is also the largest extant land artiodactyl. Despite their physical resemblance to and other terrestrial even-toed ungulates, the closest living relatives of the hippopotamids are (, , , etc.), from which they diverged about 55 million years ago. Hippos are recognisable for their barrel-shaped torsos, wide-opening mouths with large Canine tooth tusks, nearly hairless bodies, short legs, and large size: adults average for bulls (males) and for cows (females).
Hippos inhabit rivers, lakes, and mangrove swamps. Territorial bulls each preside over a stretch of water and a group of five to thirty cows and calves. Mating and birth both occur in the water. During the day, hippos remain cool by staying in water or mud, emerging at dusk to graze on grasses. While hippos rest near each other in the water, grazing is a solitary activity and hippos typically do not display territorial behaviour on land. Hippos are among the most dangerous animals in the world due to their aggressive and unpredictable nature. They are threatened by habitat loss and poaching for their meat and ivory (canine teeth).
Five subspecies of hippos have been described based on morphological differences in their skulls as well as differences in geographical range:
The most recent theory of the origins of Hippopotamidae suggests hippos and whales shared a common semiaquatic ancestor that branched off from other artiodactyls around . This hypothesised ancestral group likely split into two branches again around .
One branch would evolve into cetaceans, possibly beginning about , with the protowhale Pakicetus and other early whale ancestors collectively known as Archaeoceti. This group eventually underwent aquatic adaptation into the completely aquatic . The other branch became the anthracotheres, a large family of four-legged beasts, the earliest of which in the late Eocene would have resembled skinny hippos with comparatively smaller, narrower heads. All branches of the anthracotheres, except that which evolved into Hippopotamidae, became extinct during the Pliocene, leaving no descendants.
A rough evolutionary lineage of the hippo can thus be traced from Eocene and Oligocene species: from Anthracotherium and Elomeryx to the Miocene species Merycopotamus and Libycosaurus and finally the very latest anthracotheres in the Pliocene. These groups lived across Eurasia and Africa. The discovery of Epirigenys in East Africa, which was likely a descent of Asian anthracotheres and a sister taxon to Hippopotamidae, suggests that hippo ancestors entered Africa from Asia around . An early hippopotamid is the genus Kenyapotamus, which lived in Africa from 15 to . Hippopotamid species would spread across Africa and Eurasia, including the modern pygmy hippo. From 7.5 to , a possible ancestor to the modern hippo, Archaeopotamus, lived in Africa and the Middle East. The oldest records of the genus Hippopotamus date to the Pliocene (5.3–2.6 million years ago). The oldest unambiguous records of the modern H. amphibius date to the Middle Pleistocene, though there are possible Early Pleistocene records.
Hippopotamus gorgops from the Early Pleistocene to the early Middle Pleistocene of Africa and West Asia grew considerably larger than the living hippopotamus, with an estimated body mass of over .van der Made J, Sahnouni M & Boulaghraief K. 2017. Hippopotamus gorgops from El Kherba (Algeria) and the context of its biogeography. In Proceedings of the II Meeting of African Prehistory: Burgos 15–16 April 2015, Sahnouni M, Semaw S, Rios Garaizar J (eds). CENIEH: Burgos; 135–169.Chaix L, Faure M, Guérin C, Honegger M. Kaddanarti, a Lower Pleistocene Assemblage from Northern Sudan. In: Krzyżaniak L, Kroeper K, Kobusiewicz M, editors. Recent Research into the Stone Age of Northeastern Africa. Poznań: Poznań Archaeological Museum; 2000. p. 33–46. Hippopotamus antiquus ranged throughout Europe, extending as far north as Britain during the Early and Middle Pleistocene epochs, before being replaced by the modern H. amphibius in Europe during the latter part of the Middle Pleistocene. The Pleistocene also saw a number of dwarf species evolve on several Mediterranean islands, including Crete ( Hippopotamus creutzburgi), Cyprus (the Cyprus dwarf hippopotamus, Hippopotamus minor), Malta ( Hippopotamus melitensis), and Sicily ( Hippopotamus pentlandi). Of these, the Cyprus dwarf hippo survived until the end of the Pleistocene or early Holocene. Evidence from the archaeological site Aetokremnos continues to cause debate on whether or not the species was driven to extinction or even encountered by humans.
Hippos have barrel-shaped bodies with short tails and legs, and an hourglass-shaped skull with a long snout. Their skeletal structures are , adapted to carrying their enormous weight, and their dense bones and low centre of gravity allows them to sink and move along the bottom of the water. Hippopotamuses have small legs (relative to other megafauna) because the water in which they live reduces the weight burden.
The toes are webbed and the pelvis rests at an angle of 45 degrees. Though chubby-looking, hippos have little fat. The eyes, ears, and nostrils of hippos are placed high on the roof of their skulls. This allows these organs to remain above the surface while the rest of the body is submerged. The nostrils and ears can close when underwater while nictitating membranes cover the eyes. The of the hippo are more horizontally positioned, much like baleen whales. Underneath are throat tissues, where vibrations are transmitted to produce underwater calls.The hippo's jaw powered by huge masseter muscle and ,
and the hinge is located far back enough so that they can open their mouths at 100–110 degrees. Extensions at the back of the jaw create more surface area for muscle attachment, which gives them large, droopy cheeks This allows them to achieve their gape without tearing any tissue. On the lower jaw, the and Canine tooth grow continuously, the former reaching , while the latter can grow to up to . The lower canines are sharpened through contact with the smaller upper canines. The canines and incisors are used mainly for combat instead of feeding, and the jaws are too rigid for side to side motion, making them less efficient for chewing. Hippos rely on their flattened, horny lips to grasp and pull grasses which are then passed to the molars, which have complex Tooth enamel folds on their chewing surface. The hippo is considered to be a pseudoruminant; it has a complex three-chambered stomach, but does not "chew cud".Hippo skin is thick across much of its body with little hair. The animal is mostly purplish-grey or blue-black, but brownish-pink on the underside and around the eyes and ears. Their skin secretes a natural, red-coloured sunscreen substance that is sometimes referred to as "blood sweat" but is neither blood nor sweat. This secretion is initially colourless and turns red-orange within minutes, eventually becoming brown. Two highly acidic pigments have been identified in the secretions; one red hipposudoric acid and one orange norhipposudoric acid, which inhibit the growth of disease-causing bacteria and their light-absorption profile peaks in the ultraviolet range, creating a sunscreen effect. Regardless of diet, all hippos secrete these pigments so food does not appear to be their source; rather, they may be synthesised from precursors such as the amino acid tyrosine. This natural sunscreen cannot prevent the animal's skin from cracking if it stays out of water too long.
The testes of the males do not fully descend and a scrotum is not present. In addition, the penis retracts into the body when not erection. The genitals of the female hippos are unusual in that the vagina is ridged and the vulval vestibule has two large, protruding diverticula. Both of these have an unknown function.
A hippo's lifespan is typically 40 to 50 years. Donna the Hippo was one of the oldest living hippos in captivity. She lived at the Mesker Park Zoo in Evansville, Indiana, in the US until her death in 2012 at the age of 61. Two hippos share the record for the oldest hippo ever recorded at 65 years. Bertha, a female Hippo, who lived in the Manila Zoo in the Philippines since it first opened in 1959 until her death in July 2017, and male Hippo Lu, from the Ellie Schiller Homosassa Springs Wildlife State Park, who was born at the San Diego Zoo on January 26th, 1960, and died at Homosassa, where he died in June 2025, also at the age of 65. Two other notably long-lived hippos were the female Tanga (19341995) at the Hellabrunn Zoo in Munich, Germany and male Blackie at the Cleveland Metroparks Zoo (19532014). Currently, the oldest hippo in captivity is believed to be Mae Mali at the Khao Kheow Open Zoo in Thailand, who will turn 60 in September 2025.
Archaeological evidence exists of its presence in the Levant, dating to less than 3,000 years ago. The species was common in Egypt's Nile region during antiquity, but it has since been driven out. According to Pliny the Elder, in his time, the best location in Egypt for capturing this animal was in the Saite nome;
( English translation; Latin original) the animal could still be found along the Damietta branch of the Nile after the Arab conquest in 639. Reports of the slaughter of the last hippo in Natal Province were made at the end of the 19th century. Hippos are still found in the rivers and lakes of the northern Democratic Republic of the Congo, Uganda, Tanzania, and Kenya, north through to Ethiopia, Somalia, and Sudan, west to The Gambia, and south to South Africa.Genetic evidence suggests common hippos in Africa experienced a marked population expansion during or after the Pleistocene, attributed to an increase in water bodies at the end of the era. These findings have important conservation implications, as hippo populations across the continent are currently threatened by loss of access to fresh water. Hippos are also subject to unregulated hunting and poaching. The species is included in Appendix II of the Convention on International Trade in Endangered Species (CITES) meaning international export/import (including in parts and derivatives) requires CITES documentation to be obtained and presented to border authorities.
As of 2017, the IUCN Red List drawn up by the International Union for Conservation of Nature (IUCN) lists the species as vulnerable, with a stable population estimated between 115,000 and 130,000 animals. The hippo population has declined most dramatically in the Democratic Republic of the Congo. By 2005, the population in Virunga National Park had dropped to 800 or 900 from around 29,000 in the mid-1970s. This decline is attributed to the disruptions caused by the Second Congo War. The poachers are believed to be Mai-Mai rebels, underpaid Congolese soldiers, and local militia groups. Reasons for poaching include the belief hippos are harmful to society, as well as financial gain. As of 2016, the Virunga hippo population appears to have increased again, possibly due to better protection from park rangers, who have worked with local fishermen. The sale of hippo meat is illegal, but black-market sales are difficult for Virunga National Park officers to track. Hippo meat is highly valued in some areas of central Africa and the teeth may be used as a replacement for elephant ivory.
A population of hippos exists in Colombia, descended from captive individuals that escaped from Pablo Escobar's estate after his death in 1993. Their numbers grew to 100 by the 2020s and ecologists believe the population should be eradicated, as they are breeding rapidly and are an increasing menace to humans and the environment. Attempts to control them include sterilisation and culling.
Despite being semiaquatic, an adult hippo is not a particularly good swimmer, nor can it float. It rarely enters deep water; when it does, the animal moves by bouncing off the bottom. An adult hippo surfaces every four to six minutes, while young need to breathe every two to three minutes. Hippos move on land by , and limb movements do not change between speeds. They can reach an airborne stage (a stage when all limb are off the ground) when they move fast enough. Hippos are reported to reach but this has not been confirmed. They are incapable of jumping but can walk up steep banks. The hippopotamus sleeps with both hemispheres of the brain resting, as in all land mammals, and usually sleeps on land or in water with the nostrils exposed. Despite this, it may be capable of sleeping while submerged, intermittently surfacing to breathe without waking. They appear to transition between different phases of sleep more quickly than other mammals.
Because of their size and their habit of taking the same paths to feed, hippos can have a significant impact on the land across which they walk, keeping the land clear of vegetation and depressing the ground. Over prolonged periods, hippos can divert the paths of swamps and channels. By defecating in the water, the animals also appear to pass on microbes from their gut, affecting the biogeochemical cycle. On occasion, hippos have been filmed eating carrion, usually near the water. There are other reports of meat-eating and even cannibalism and predation. Hippos' stomach anatomy lacks adaptions to carnivory and meat-eating is likely caused by lack of nutrients or just an abnormal behaviour.
Hippos engage in "muck-spreading" which involves defecating while spinning their tails to distribute the faeces over a greater area. Muck-spreading occurs both on land and in water and its function is not well understood. It is unlikely to serve a territorial function, as the animals only establish territories in the water. They may be used as trails between the water and grazing areas. "Yawning" serves as a threat display. When fighting, bulls use their incisors to block each other's attacks and their large canines as offensive weapons. When hippos become over-populated or a habitat shrinks, bulls sometimes attempt infanticide, but this behaviour is not common under normal conditions.
The most common hippo vocalisation is the "wheeze honk", which can travel over long distances in air. This call starts as a high-pitched squeal followed by a deeper, resonant call. The animals can recognise the calls of other individuals. Hippos are more likely to react to the wheeze honks of strangers than to those they are more familiar with. When threatened or alarmed, they produce exhalations, and fighting bulls will bellow loudly. Hippos are recorded to produce clicks underwater which may have echolocative properties. They have the unique ability to hold their heads partially above the water and send out a cry that travels through both water and air; individuals respond both above and below water.
Mother hippos are very protective of their young, not allowing others to get too close. One cow was recorded protecting a calf's carcass after it had died. Calves may be temporarily kept in nurseries, guarded by one or more adults, and will play amongst themselves. Like many other large mammals, hippos are described as k-selection, in this case typically producing just one large, well-developed infant every couple of years (rather than many small, poorly developed young several times per year, as is common among small mammals such as rodents). Calves no longer need to suckle when they are a year old.
Hippos occasionally visit cleaning stations in order to be cleaned of parasites by certain species of fishes. They signal their readiness for this service by opening their mouths wide. This is an example of mutualism, in which the hippo benefits from the cleaning while the fish receive food.
Hippo defecation creates allochthonous deposits of organic matter along the river beds. These deposits have an unclear ecological function. A 2015 study concluded hippo dung provides nutrients from terrestrial material for fish and aquatic invertebrates, while a 2018 study found that their dung can be toxic to aquatic life in large quantities, due to absorption of dissolved oxygen in water bodies.The parasitic flatworm Oculotrema hippopotami infests hippopotamus eyes, mainly the nictitating membrane. It is the only monogenean species (which normally live on fish) documented to live on a mammal.
The hippo was also known to the Ancient Greece and Ancient Rome. The Greek historian Herodotus described the hippo in The Histories (written 440 BC) and the Roman naturalist Pliny the Elder wrote about the hippo in his encyclopedia Naturalis Historia (written circa 77 AD).
The Yoruba people called the hippo erinmi, which means "elephant of the water". Some individual hippos have achieved international fame. Huberta became a celebrity during the Great Depression for trekking a great distance across South Africa.
According to the Ptolemaic historian Manetho, the pharaoh Menes was carried off and then killed by a hippopotamus.
Hippos generally breed well in captivity; birth rates are lower than in the wild, but this can be attributed to zoos' desire to limit births, since hippos are relatively expensive to maintain. Starting in 2015, the Cincinnati Zoo built a US$73 million exhibit to house three adult hippos, featuring a tank. Modern hippo enclosures also have a complex filtration system for waste, an underwater viewing area for the visitors, and glass that may be up to thick and capable of holding water under pressures of .
Hippos have been the subjects of various African folklore. According to a San people story, when the Creator deity assigned each animal its place in nature, the hippos wanted to live in the water, but were refused out of fear they might eat all the fish. After begging and pleading, the hippos were finally allowed to live in the water on the condition they would eat grass instead of fish, and fling their dung so it can be inspected for fish bones. In a Ndebele tale, the hippo originally had long, beautiful hair, but it was set on fire by a jealous hare and the hippo had to jump into a nearby pool. The hippo lost most of his hair and was too embarrassed to leave the water.
Hippopotamuses were rarely depicted in European art during the Renaissance and Baroque periods, due to less access to specimens by Europeans. One notable exception is Peter Paul Rubens' The Hippopotamus and Crocodile Hunt (1615–1616). Ever since Obaysch inspired the "Hippopotamus Polka", hippos have been popular animals in Western culture for their rotund appearance, which many consider comical. The Disney film Fantasia featured a ballerina hippo dancing to the opera La Gioconda. The film Hugo the Hippo is set in Tanzania and involves the title character trying to escape being slaughtered with the help of local children. The Madagascar films feature a hippo named Gloria.
Hippos even inspired a popular board game, Hungry Hungry Hippos.Among the most famous poems about the hippo is "The Hippopotamus" by T. S. Eliot, where he uses a hippo to represent the Catholic Church. Hippos are mentioned in the novelty Christmas song "I Want a Hippopotamus for Christmas" that became a hit for child star Gayla Peevey in 1953. They also featured in the popular "The Hippopotamous Song" by Flanders and Swann.
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