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File:Berberis thunbergii berries.jpgJapanese barberries File:Norwegian blueberry.jpg File:Ribes rubrum2005-07-17.JPG File:Lonicera coerulea a3.jpgHoneysuckle File:Stachelbeere (Ribes uva-crispa).jpg File:Rubus chamaemorus close-up.JPGCloudberry File:Vaccinium corymbosum a2.jpgHighbush blueberries File:Black Butte blackberry.jpg
A berry is a small, pulpy, and often edible . Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a or pit although many or seeds may be present. Common examples of berries in the culinary sense are , , , , , , and . In Britain, soft fruit is a term for such fruits.

The common usage of the term "berry" is different from the scientific or botanical definition of a berry, which refers to a produced from the ovary of a single flower where the outer layer of the ovary wall develops into an edible fleshy portion(). The botanical definition includes many fruits that are not commonly known or referred to as berries,

(2026). 9780143419549, DK Pub..
such as , , , , , and . Fruits commonly considered berries but excluded by the botanical definition include strawberries, raspberries, and , which are , and mulberries, which are . and are giant berries that fall into the category "pepos". A plant bearing berries is said to be or .

Berries are eaten worldwide and often used in , , , or . Some berries are commercially important. The berry industry varies from country to country as do types of berries cultivated or growing in the wild. Some berries such as raspberries and strawberries have been bred for hundreds of years and are distinct from their wild counterparts, while other berries, such as and , grow almost exclusively in the wild.

While many berries are edible, some are to humans, such as those of deadly nightshade and . Others, such as the , , and , are poisonous when , but are edible when ripe.


History
Berries have been valuable as a food source for humans since before the start of agriculture, and remain a food source for other . They were a seasonal staple for early for thousands of years, and wild berry gathering remains a popular activity in Europe and North America today. In time, humans learned to store berries so that they could be used in the winter. They may be made into , and among Native Americans, mixed with meat and fats as .
(2026). 9780521402156, Cambridge University Press. .

Berries also began to be cultivated in Europe and other countries. Some species of blackberries and raspberries of the genus have been cultivated since the 17thcentury, while smooth-skinned blueberries and cranberries of the genus have been cultivated in the United States for over a century. In Japan, between the 10th and 18thcenturies, the terms and (: ; : ) referred to many berry crops. The most widely cultivated berry of modern times is the , which is produced globally at twice the amount of all other berry crops combined.

The strawberry was mentioned by ancient Romans, who thought it had medicinal properties,

(2026). 9781423608813, . .
but it was then not a staple of agriculture. Woodland strawberries began to be grown in French gardens in the 14thcentury. The ( F. moschata), also known as the hautbois strawberry, began to be grown in European gardens in the late 16thcentury. Later, the Virginia strawberry was grown in Europe and the United States. The most commonly consumed strawberry, the garden strawberry( F. ananassa), is an accidental hybrid of the Virginia strawberry and a Chilean variety Fragaria chiloensis. It was first noted by a French gardener around the mid 18thcentury that, when F. moschata and F. virginiana were planted in between rows of F. chiloensis, the Chilean strawberry would bear abundant and unusually large fruits. Soon after, began to study the breeding of strawberries and made several discoveries crucial to the science of plant breeding, such as the sexual reproduction of strawberry. Later, in the early 1800s, English breeders of strawberry made varieties of F. ananassa which were important in strawberry breeding in Europe, and hundreds of have since been produced through the breeding of strawberries.
(2026). 9783642160578, Springer. .


Etymology
The word () comes from Proto-Germanic, variously reconstructed as , , basjom (source also of , , , , , , ), which is of unknown origin. This and "" are the only fruit names in modern English which are descended from "native" Germanic words.


Botanical definition
In terminology, a berry is a with and pulp produced from the ovary of a single flower. It is fleshy throughout, except for the seeds. It does not have a special "line of weakness" along which it splits to release the seeds when ripe (i.e. it is indehiscent). A berry may develop from an ovary with one or more (the female reproductive structures of a flower). The seeds are usually embedded in the fleshy interior of the ovary, but there are some non-fleshy examples such as , with air rather than pulp around their seeds. The differences between the everyday and botanical uses of "berry" result in three categories: those fruits that are berries under both definitions; those fruits that are botanical berries but not commonly known as berries; and those parts of plants commonly known as berries that are not botanical berries, and may not even be fruits.

Berries under both definitions include , , , and the fruits of many other members of the , as well as , and . The fruits of some "currants" ( species), such as , and , are botanical berries, and are treated as horticultural berries (or as soft fruit in the UK), even though their most commonly used names do not include the word "berry".

Botanical berries not commonly known as berries include , , , (aubergines), , , and .

There are several different kinds of fruits which are commonly called berries, but are not botanical berries. , , and are kinds of ; they contain seeds from different ovaries of a single flower. In aggregate fruits like blackberries, the individual "fruitlets" making up the fruit can be clearly seen. The fruits of may be called "sloe berries",

(2003). 9780306477065, Springer Science & Business Media. .
but botanically are small or drupes, like or .

and are commonly said to have berries, but these plants do not produce botanical fruits at all: they are , specifically , not (flowering plants). Their "berries" are highly-modified seed-bearing . In , used to flavour , the cone scales, which are hard and woody in most conifers, are instead soft and fleshy when ripe. The bright red berries of yews consist of a fleshy outgrowth() almost enclosing the poisonous seed. The resemblance of these plant structures to botanical berries provides a striking example of convergent evolution in different plant .


Cultivation
have been grown in gardens in Europe since the 14thcentury. were domesticated starting in 1911, with the first commercial crop in 1916. of all varieties are not fully domesticated, but domestication was attempted from 1994 to 2010 for the economically significant western huckleberry. Many other varieties of are likewise not domesticated, with some being of commercial importance.


Agricultural methods
Like most other food crops, berries are commercially grown, with both conventional pest management and integrated pest management(IPM) practices. Organically certified berries are becoming more widely available.:5

Many soft fruit berries require a period of temperatures between for breaking dormancy. In general, require 200–300hours, 650–850hours, 700hours, 800–1700hours, and 800–1500hours, and 2000hours. However, too low a temperature will kill the crops: blueberries do not tolerate temperatures below , raspberries, depending on variety, may tolerate as low as , and blackberries are injured below . Spring frosts are, however, much more damaging to berry crops than low winter temperatures. Sites with moderate slopes(), facing north or east in the Northern Hemisphere, near large bodies of water, which regulate spring temperature, are considered ideal in preventing spring frost injury to the new leaves and flowers. All berry crops have shallow . Many US land-grant university extension offices suggest that strawberries should not be planted more than five years on the same site, due to the danger of black root rot (though many other illnesses go by the same name), which in the past has been controlled in major commercial production by annual fumigation but which is largely prohibited now. Besides the number of years in production, , the frequency of fumigation, and herbicide usages increase the appearance of black root rot in strawberries. Raspberries, blackberries, strawberries, and many other berries are susceptible to verticillium wilt. Blueberries and cranberries grow poorly if the clay or silt content of the soil is more than 20%, while most other berries tolerate a wide range of . For most berry crops, the ideal soil is well drained sandy , with of 6.2–6.8 and a moderate to high organic content; however, blueberries have an ideal pH of 4.2–4.8 and can be grown on muck soils, while blueberries and cranberries prefer poorer soils with lower cation exchange, lower calcium, and lower levels of phosphorus.

Growing most berries organically requires the use of proper , the right mix of , and the cultivation of the correct in the soil. As blueberries and cranberries thrive in soils that are not hospitable to most other plants, and conventional fertilizers are toxic to them, the primary concern when growing them organically is bird management.

Postharvest small fruit berries are generally stored at relative humidity and . Cranberries, however, are frost sensitive, and should be stored at . Blueberries are the only berries that respond to , but flavor does not improve after harvest, so they require the same treatment as other berries. Removal of ethylene may reduce disease and spoilage in all berries. Precooling within one to two hours post-harvest to storage temperature, generally, via forced air cooling increases the storage life of berries by about a third. Under optimum storage conditions, raspberries and blackberries last for two to five days, strawberries 7–10days, blueberries two to four weeks, and cranberries two to four months. Berries can be shipped under high carbon dioxide or modified atmosphere of carbon dioxide for high carbon dioxide or carbon dioxide and oxygen for a modified atmosphere container to increase shelf life and prevent grey mold rot.


Breeding
Several discoveries in the science of breeding berries were made in the 18thcentury by in his work on strawberries. In the traditional technique of , berries with specific desirable characteristics were chosen and allowed to sexually reproduce with other berries, and offspring with improved traits could then be selected and used for further crossing. Plants may be with different species within the same ; hybridization between different genus may also be possible, but more difficult. Breeding may seek to increase the size and yield of the fruit, improve the flavor and quality of its nutrient content, such as antioxidants, expand the harvest season, and produce with resistance to diseases, tolerance of hot or cold conditions, and other desirable traits.
(2011). 9781439856604, . .
Advancements in molecular biology and genetic engineering allow for a more efficient and better targeted approach in the selection for a desirable , via marker-assisted selection, for example. Genetic modification techniques can also be used for breeding berries.


Horticultural soft fruit berries
Some fruit not commonly referred to as berries and not always botanically berries are included by US land-grant university extension offices in their guides for berry cultivation, or in guides for identifying local wild edible and non-edible berries. Examples include , American persimmons, , Pacific crabapples, and .


Commercial production
In the year 2005, there were of land worldwide cultivating berries, with produced.
(2007). 9781420006148, CRC Press. .
:4


Economics
In certain regions, berrypicking can be a large part of the economy, and it is becoming increasingly common for countries such as Sweden and Finland to from Thailand or Bulgaria to do berry picking. This practice has come under scrutiny in the past years because of the low wages and poor living standard for the "berry-pickers", as well as the lack of worker safety.

In the late 2010s in the US, reduced migration from Mexico and Central America and increased minimum wage standards have made finding "stoop-work" labourers to pick the strawberry crop difficult and costly.


Phytochemicals and color
Once , berries have a contrasting color to their background (often of green leaves), making them visible and attractive to animals and birds.
(2026). 9780226471051, University of Chicago Press. .
This assists the wide dispersal of the plants' seeds.

Berry colors are due to natural , including , such as , together with other localized mainly in berry skins, seeds and leaves. Although berry pigments have properties , there is no evidence established to date that berry pigments have actual antioxidant or any other functions within the human body. Consequently, it is not permitted to claim that foods containing polyphenols have antioxidant health value on product labels in the United States or Europe. Guidance for Industry, Food Labeling; Nutrient Content Claims; Definition for "High Potency" and Definition for "Antioxidant" for Use in Nutrient Content Claims for Dietary Supplements and Conventional Foods U.S. Department of Health and Human Services, Food and Drug Administration, Center for Food Safety and Applied Nutrition, June 2008


Culinary significance

Use in baked goods
Berries are commonly used in pies or , such as , , and .

Berries are often used in baking, such as blueberry muffins, blackberry muffins, , berry crisps, berry cakes, berry buckles, berry , berry , and berry cookies. Berries are commonly incorporated whole into the batter for baking, and care is often taken so as to not burst the berries. Frozen or dried berries may be preferable for some baked berry products. Fresh berries are also often incorporated into baked berry desserts, sometimes with , either as a filling to the dessert or as a topping.


Beverages
Berries are often added to water and/or , as in , which accounts for 95% of cranberry crop usage, juice, , juice, juice, juice, and juice. is the principal alcoholic beverage made from berries(grapes). are commonly made out of other berries. In most cases, sugars must be added to the berry juices in the process of to increase the alcohol content of the wine. Examples of fruit wines made from berries include: elderberry wine, strawberry wine, blueberry wine, blackberry wine, redcurrant wine, huckleberry wine, goji wine and cranberry wine. Berries are used in some styles of beer, particularly (made with raspberry) and other fruit .


Dried
, and sultanas are examples of dried grape berries, and many other commercially important berries are available in dried form.


Fruit preserves
Berries are perishable fruits with a short shelf life, and are often preserved by , , or making . Berries such as , , , , ,
(2026). 9781317846055, Taylor & Francis. .
, and are often used in jams and jellies. In the United States, Native Americans were "the first to make preserves from blueberries".


Other usages
Chefs have created quick pickled soft fruit, such as blackberries, strawberries, and blueberries. Strawberries can be battered and quickly fried in a . Sauces made from berries, such as cranberry sauce, can be frozen until hard, battered, and deep fried. Cranberry sauce is a traditional food item for , and similar sauces can be made from many other berries such as blueberries, raspberries, blackberries, and huckleberries.


In culture

Dye
Berries have been used in some cultures for . Many berries contain juices that can easily stain, affording use as a natural dye. For example, blackberries are useful for making dyes, especially when ripe berries can easily release juice to produce a effect. Rubus berries, such as blackberry, raspberry, , , , and all produce dye colors. These were once used by Native Americans. In Hawaii, the native Hawaiian raspberry called was used to dye with lavender and pink hues, whereas berries from the were used for blue coloration, and berries from the were used to produce green coloration.


Research
Berry consumption is under preliminary research for the potential to improve nutrition and affect chronic diseases. A 2016 found that berry consumption can significantly lower body mass index, low density lipoprotein (LDL) and systolic blood pressure.


See also
  • List of berries


Further reading


External links
  • of the National Berry Crops Initiative (United States)

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