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Storks are large, long-legged, long-necked with long, stout Beak. They belong to the family Ciconiidae, and make up the order Ciconiiformes . Ciconiiformes previously included a number of other families, such as and , but those families have been moved to other orders.
Storks dwell in many regions and tend to live in drier habitats than the closely related , and ; they also lack the powder down that those groups use to clean off fish slime. Bill-clattering is an important mode of communication at the nest. Many species are bird migration. Most storks eat , fish, , , small and small . There are 20 living species of storks in six genera.
Various terms are used to refer to groups of storks, About the Wood Stork: Denizens of the Wetlands , Accessed on 13.12.2010 two frequently used ones being a muster of storks and a phalanx of storks.
Storks tend to use soaring, gliding flight, which conserves energy. Soaring requires thermal air currents. Ottomar Anschütz's famous 1884 album of photographs of storks inspired the design of Otto Lilienthal's experimental Glider aircraft of the late nineteenth century. Storks are heavy, with wide : the marabou stork, with a wingspan of and weight up to , joins the Andean condor in having the widest wingspan of all living land birds.
Their bird nest are often very large and may be used for many years. Some nests have been known to grow to over in diameter and about in depth. All storks were once thought to be monogamous, but this is only partially true. While storks are generally socially monogamous, some species exhibit regular extra-pair breeding.
Popular conceptions of storks' fidelity, serial monogamy, and doting parental care contribute to their prominence in mythology and culture, especially in western folklore as the deliverers of newborn humans.
All 20 stork species have been assessed by the IUCN and carry a confident Red List status. However, the assessment for several species were based on incorrect assumptions and a general absence of sound information on stork habits.
Storks were distinct and possibly widespread by the Oligocene. Like most families of aquatic birds, storks seem to have arisen in the Paleogene, maybe 40–50 million years ago (mya). For the fossil record of living genera, documented since the Middle Miocene (about 15 mya) at least in some cases, see the genus articles.
No species or subspecies of stork is known to have gone extinct in historic times. A systematic literature review uncovered nearly 1,000 papers on storks, but showed most stork species to lack scientific understanding suggesting that many species should be classified as Data Deficient on the IUCN Red List. A Ciconia bone found in a rock shelter on the island of Réunion was probably of a bird taken there as food by early settlers; no known account mentions the presence of storks on the Mascarene Islands.
The fossil genera Eociconia (Middle Eocene of China) and Ciconiopsis (Deseado Early Oligocene of Patagonia, Argentina) are often tentatively placed with this family. A "ciconiiform" fossil fragment from the Touro Passo Formation found at Arroio Touro Passo (Rio Grande do Sul, Brazil) might be of the living wood stork M. americana; it is at most of Late Pleistocene age, a few 10,000s of years.Schmaltz Hsou, Annie (2007). O estado atual do registro fóssil de répteis e aves no Pleistoceno do Estado do Rio Grande do Sul, Brasil "The. Talk held on 2007-JUN-20 at Quaternário do RS: integrando conhecimento, Canoas, Rio Grande do Sul, Brazil. PDF abstract
The beak of storks are large to very large, and vary considerably between the genera. The shape of the bills is linked to the diet of the different species. The large bills of the Ciconia storks are the least specialized. Larger are the massive and slightly upturned bills of the Ephippiorhynchus and the jabiru. These have evolved to hunt for fish in shallow water. Larger still are the massive daggers of the two adjutants and marabou ( Leptoptilos), which are used to feed on carrion and in defense against other scavengers, as well as for taking other prey. The long, ibis-like downcurved bills of the Mycteria storks have sensitive tips that allow them to detect prey by touch (tactilocation) where cloudy conditions would not allow them to see it. The most specialised bills of any storks are those of the two openbills ( Anastomus), which as their name suggests, is open in the middle when their bill is closed. These bills have evolved to help openbills feed on their primary prey item, aquatic snails.
Although it is sometimes reported that storks lack syrinxes and are mute, they do have syrinxes, and are capable of making some sounds, although they do not do so often. The syrinxes of storks are "variably degenerate" however, and the syringeal membranes of some species are found between tracheal rings or cartilage, an unusual arrangement shared with the ovenbirds.
Storks are more diverse and common in the tropics, and the species that live in temperate climates for the most part bird migration to avoid the worst of winter. They are fairly diverse in their habitat requirements. Some species, particularly the Mycteria "wood storks" and Anastomus openbills, are highly dependent on water and aquatic prey, but many other species are far less dependent on this habitat type, although they will frequently make use of it. Species like the Marabou stork and Abdim's stork will frequently be found foraging in open grasslands of savannah. Preferred habitats include flooded grasslands, light woodland, marshes and paddyfields, , river backwaters and ponds. Many species will select shallow pools, particularly when lakes or rivers are drying out, as they concentrate prey and make it harder for prey to escape, or when monsoonal rainfall increases water depth of larger waterbodies. Some species like the woolly-necked storks and lesser adjutant storks have adapted to changing crops of tropical agricultural landscapes that enables them to remain resident despite the transformations brought about by seasonal crops. In South Africa, the woolly-necked storks have adapted to artificial feeding and now largely nest on trees in gardens with swimming pools.
Less typical habitats include the dense temperate forests used by European Black stork, or the rainforest habitat sought by Storm's stork in South East Asia. They generally avoid marine habitats, with the exception of the lesser adjutant, milky stork and wood stork, all of which forage in mangroves, lagoons and estuarine mudflats. A number of species, especially woolly-necked storks, black-necked storks, Asian openbills and lesser adjutant Storks in south Asia, have adapted to highly modified human habitats, for foraging and breeding. In the absence of persecution several stork species breed close to people, and species such as the marabou, greater adjutant, and white stork feed at landfill sites.
The Abdim's stork is another migrant, albeit one that migrates within the tropics. It breeds in northern Africa, from Senegal to the Red Sea, during the wet season, and then migrates to Southern Africa. Many species that are not regular migrants will still make smaller movements if circumstances require it; others may migrate over part of their range. This can also include regular commutes from nesting sites to feeding areas. Wood storks have been observed feeding from their breeding colony.
Openbills are specialists in freshwater molluscs, particularly apple snails. They feed in small groups, and sometimes African openbills ride on the backs of hippos while foraging. Having caught a snail it will return to land or at least to the shallows to eat it. The fine tip of the bill of the openbills is used to open the snail, and the saliva has a narcotic effect, which causes the snail to relax and simplifies the process of extraction.
The other genera of storks are more generalised. Ciconia storks are very generalised in their diets, and some species including Abdim's stork and marabous will feed in large flocks on swarms of locusts and at wildfires. This is why white storks and Abdim's storks are known as "grasshopper birds". Ephippiorhynchus are carnivorous though have a very diverse diet when living on human modified habitats such as agricultural landscapes. The foraging method used by the generalists is to stalk or walk across grassland or shallow water, watching for prey.
Greek mythology and Roman mythology portrays storks as models of parental devotion. The 3rd century Roman writer Aelian, citing the authority of Alexander of Myndus, noted in his De natura animalium (book 3, chapter 23) that aged storks flew away to oceanic islands where they were transformed into humans as a reward for their piety towards their parents. Storks were also thought to care for their aged parents, feeding them and even transporting them, and children's books depicted them as a model of filial values. A Greek law called Pelargonia, from the Ancient Greek word pelargos for stork, required citizens to take care of their aged parents. The Greeks also held that killing a stork could be punished with death.
Storks feature in several of Aesop's Fables, most notably in The Farmer and the Stork, The Fox and the Stork, and The Frogs Who Desired a King. The first fable involves a stork who is caught with a group of cranes who are eating grain in a farmer's field, with the moral that those who associate with wicked people can be held accountable for their crimes. The Fox and the Stork involves a fox who invites a stork for dinner and provides soup in a dish that the stork cannot drink from, and is in turn invited for dinner by the stork and given food in a narrow jug which he cannot access. It cautions readers to follow the principle of do no harm. The third fable involves a group of frogs that are dissatisfied with the king that Zeus has given them, an inanimate log, and who are then punished with a new (a water-snake in some versions) who eats the frogs. King Stork has subsequently entered the English language as a term for a particularly tyrannical ruler.
Famous is the role that the fable played in historical development of psychoanalysis: the name ‘chimney sweeping’, which the first of all patients gave to her talking cure, is a free association with the place through which the bird used to bring babies into house. Psychoanalyst Marvin Margolis suggests the enduring nature of the stork fable of the newborn is linked to its addressing a psychological need, in that it allays the discomfort of discussing sex and procreation with children. Birds have long been associated with the maternal symbols from pagan goddesses such as Juno, to the Holy Ghost, and the stork may have been chosen for its white plumage (depicting purity), size, and flight at high altitude (likened to flying between Earth and Heaven).
There were negative aspects to stork folklore as well; a Polish folktale relates how God made the stork's plumage white, while the Devil gave it black wings, imbuing it with both good and evil impulses. They were also associated with handicapped or stillborn babies in Germany, explained as the stork having dropped the baby en route to the household, or as revenge or punishment for past wrongdoing. A mother who was confined to bed around the time of childbirth was said to have been "bitten" by the stork. In Denmark, storks were said to toss a nestling off the nest and then an egg in successive years. In medieval England, storks were also associated with adultery, possibly inspired by their courtship rituals. Their preening and posture saw them linked with the attribute of self-conceit. Children of African American slaves were sometimes told that white babies were brought by storks, while black babies were born from buzzard eggs.
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