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Ridgehead
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Ridgeheads, also known as bigscales, are a family ( Melamphaidae, from the melanos black and amphi by) of small, deep-sea . The family contains approximately 37 in five ; their distribution is worldwide, but ridgeheads are absent from the and Mediterranean Sea. Although the family is one of the most widespread and plentiful of deep-sea families, none of its members are of interest to commercial .

These fish are named for their large scales and pronounced cranial ridges, as well as for their typically dark brown to black coloration. Ridgeheads are the largest and most diverse family of their order.


Description
The melamphid body is robust, oblong, , and slightly compressed laterally. The head is large and scaleless, with its profile either bluntly rounded or with a sharp frontal angle; it is conspicuous for its prominent ridges, which are covered by thin . The head is also cavernous, being riddled by canals—similar structures are found in the (Trachichthyidae) and (Anoplogastridae) families. Sharp spines and serrations may further adorn the head in some species. The mouth is large and oblique; the long extend to or exceed the posterior margin of the , and are lined with villiform (brushlike) . The , , , and all lack teeth. The are usually fairly small, but in Poromitra megalops they may exceed 20 per cent of the head's length in diameter.

The large scales are cycloid and imbricate; they are arranged in a longitudinal series of 12–40. Usually deciduous, these scales are largest in species; in these and some other species, the scales' circuli (concentric lines) are clearly visible to the unaided eye. The (excluding the network of cranial canals) is largely absent; when present, it is reduced to 1–2 pored scales following the operculum's upper edge. The caudal peduncle is relatively long; the is forked to emarginate and possesses 3–4 procurrent spines. The single originates behind the and and contains 1–3 weak spines and 9–18 soft rays; the pectoral fins are elongate and tapered (the superior rays being longer than the inferior rays) and contain 12–16 soft rays. The pelvic fins are thoracic to subthoracic with one spine and 6–8 soft rays; the is small and set far back, with one spine and 7–11 soft rays.

The branchiostegal rays number 7–8 and the 24–31. The largest species recorded is Poromitra curilensis, a Pacific ridgehead related to the , at up to 18 centimetres standard length (SL; a measurement excluding the caudal fin). Most ridgeheads are well under 10 centimetres SL.


Life history
Ridgeheads are meso- to ; they have been at depths as shallow as to as deep as . Some species undergo diel vertical migrations of several hundred metres; that is, they remain at depths (below ca. 400 metres) during the day but rise to surface waters (10–100 metres) at night to feed and (presumably) to avoid . Ridgeheads feed primarily on , such as , , and other small pelagic . There is also a marked segregation of life stages: and juvenile ridgeheads are present within the upper 200 metres of the water column, whereas adults are typically found much farther down.

The family is sometimes termed pseudoceanic because, rather than having an even distribution in open water, ridgeheads occur in significantly higher abundances around structural oases, notably and over continental slopes. This is also true of other deep-sea pelagic groups, such as the (Myctophidae), marine hatchetfish (Sternoptychidae), and (Phosichthyidae or Photichthyidae) families. The phenomenon is explained by the likewise abundance of prey species which are also attracted to the structures.

As pelagic spawners, ridgeheads produce many tiny which are externally; the eggs and larvae (and early juveniles) are buoyant and , drifting with the currents near the surface until the juveniles have reached the point where they are strong enough to determine their own direction. Study of the spawning habits of twospine bigscales ( Scopelogadus mizolepis bispinosus) indicates spawning occurs year-round, with no definite peaks.

Although their deep-living nature protects them from predation to some degree, ridgeheads are prey to large such as ; large such as the ( Dosidicus gigas) and Sevenstar Flying Squid ( Martialia hyadesii); (family Delphinidae); and large , such as and other .


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