The phragmocone is the chambered portion of the Exoskeleton of a cephalopod. It is divided by septa into camerae.
In most and Ammonoidea, the phragmocone is a long, straight, curved, or coiled structure, in which the camerae are linked by a siphuncle which determines buoyancy by means of gas exchange.
Despite this benefit, such a large shell adds to the mass of the animal, and hence is disadvantageous in catching fast-moving prey. Some nautiloids, such as the Silurian Ascocerida, dropped the phragmocone upon maturity, presumably to increase speed and maneuverability. They thus became the early Paleozoic equivalent of . The early coleoids and Belemnoidea adopted a different approach: the phragmocone was retained but became internal and reduced. In general the shell in cephalopods tends to be vestigial or absent.
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