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In , intergradation is the way in which two distinct are connected via areas where are found that have the characteristics of both. There are two types of intergradation: primary and secondary.


Primary intergradation
This occurs in cases where two subspecies are connected via one or more intermediate populations, each of which is in turn intermediate to its adjacent populations and exhibits more or less the same amount of variability as any other population within the . Adjacent populations and subspecies are subject to cline intergradation, and in these situations it is usually taken for granted that the clines are causally related (by natural selection) to environmental gradients.Mayr E, Ashlock PD (1991). Principles of Systematic Zoology. Second edition. McGraw-Hill. .


Secondary intergradation
When contact between a geographically isolated subspecies is reestablished with the main body of the species or with another isolate subspecies, interbreeding takes place as long as the isolate has not yet evolved an effective set of isolating mechanisms. Consequently, a relatively distinct zone or belt of hybridization will develop depending on the degree of and difference that was achieved by the previously isolated .


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