Product Code Database
Hurdiidae (synonymous with the previously named Peytoiidae) is an extinct cosmopolitan family of , a group of stem-group marine , which lived during the Paleozoic Era. It is the most long-lived radiodont clade, lasting from the Cambrian period to the Devonian period.
Hurdiidae is characterized by frontal appendages with distal region composed of 5 subequal blade-like , alongside the enlarged head carapaces and tetraradial mouthpart (oral cone).
The frontal appendages of hurdiids have a distinctive morphology, with the appendage of most species bearing five equally-sized elongate blade-like ventral spines known as endites. Subsequent podomeres were reduced in size and with only small endites or none. Each podomere bore only a single endite, unlike other radiodonts, in which the endites were paired. In most species, the endites were curved medially, so that the appendages formed a basket-like structure. Some hurdiids had greater numbers of endites, with Cordaticaris bearing seven endites of equal length. Ursulinacaris is unique among hurdiids in bearing paired endites, which is likely a transitional form between the appendage of other radiodonts and that of hurdiids.
Hurdiids exhibited a wide range of body size. The smallest known hurdiid specimen, of an unnamed species, is estimated to have had a body length of , but it is not known whether this specimen was a juvenile or an adult. Aegirocassis, the largest known hurdiid, was over long, comparable in size to the largest known arthropods.
Cladogram of Hurdiidae after Moysiuk & Caron, 2025:
Species include:
Zhenghecaris was originally described as a thylacocephalan, but it was later considered a hurdiid dorsal carapace. However, its placement is questioned by some researchers, since its classification as a radiodont is largely based on the characters of Tauricornicaris.
While Schinderhannes was originally described as a stem-arthropod with characters of both euarthropods and radiodonts, this interpretation was denied and most researchers now agree that it is a hurdiid radiodont. Dryad Data There are some researchers who have questioned its classification as a hurdiid.
Huangshandongia, Liantuoia and Proboscicaris hospes may represent species of Hurdia.
Paleobiology
The majority of hurdiids appear to have been predators that fed by sifting sediment with their frontal appendages, but some members, like Aegirocassis, Pseudoangustidontus, and possibly Cambroraster, were suspension feeders.
Distribution
Hurdiids had a global distribution. The earliest known hurdiid in the fossil record is Peytoia infercambriensis, which lived during the third age of the Cambrian in what is now the country of Poland. The group increased in diversity during the Miaolingian epoch. Post-Cambrian records of the group are rare, but the group lasted into the Devonian period, with the last known taxon being the Emsian Schinderhannes bartelsi from what is now Germany.
Classification
Hurdiidae is classified within Radiodonta, a clade of stem-group arthropods. Hurdiidae is defined phylogenetically as the most inclusive clade containing Hurdia victoria but not Amplectobelua symbrachiata, Anomalocaris canadensis, or Tamisiocaris borealis. Some authors have argued that Peytoiidae, which was named by Conway Morris and Robison, 1982, has priority over Hurdiidae, and that Hurdiidae has "yet to be properly established following ICZN standards", due to its first definition not having a character-based diagnosis, and the second being published in an online-only journal without being registered in the ZooBank database.
Tauricornicaris was previously considered as a member of Hurdiidae, but fossils of it were later reinterpreted to be euarthropod tergites.
/ref>
|
|
