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Stachys is a genus of plants, one of the largest in the mint family .Harley, R. M., et al. 2004. "Labiatae". pages 167–275. In: Kubitzki, K. (editor) and J. W. Kadereit (volume editor). The Families and Genera of Vascular Plants volume VII. Springer-Verlag: Berlin; Heidelberg, Germany. Estimates of the number of vary from about 300, to about 450.Mabberley, D. J. 2008. Mabberley's Plant-Book third edition (2008). Cambridge University Press: UK. Stachys is in the subfamily and its is Stachys sylvatica. Stachys In: Index Nominum Genericorum. In: Regnum Vegetabile (see External links below). The precise extent of the genus and its relationship to other genera in the subfamily are poorly known.


Range and naming
The distribution of the genus covers , , , and . include hedgenettle, heal-all, self-heal, woundwort, betony, Monty plant and lamb's ears. Wood betony, S. officinalis, was the most important to the of early England, and was used for many medicinal purposes from Ancient Roman times to the Early Modern period.

Stachys was by in Species Plantarum in 1753.Carolus Linnaeus. 1753. Species Plantarum 2:580. Laurentii Salvii. (see External Links below). The name is from the Greek word σταχυς ( stachys), meaning "an ear of grain",Umberto Quattrocchi. 2000. CRC World Dictionary of Plant Names volume I, page 91. CRC Press: Boca Raton; New York; Washington, DC;, USA. London, UK. (set). (see External links below). and refers to the fact that the is often a . The name woundwort derives from the past use of certain species in for the treatment of wounds.


Human uses
The Chinese artichoke or Crosne ( ), is grown for its edible . Several species are as . Woolly betony ( S. byzantina) is a popular decorative garden plant. Wood betony ( S. officinalis) was historically a highly valued medicinal plant.


Use by other species
Stachys species are used as food plants by the of some species, including the Coleophora auricella, C. lineolea, and C. wockeella, all recorded on S. officinalis. They are also widely used by the European wool carder bee ( Anthidium manicatum), which scrape the hairs from the plant in order to use them for building their nests.


Description
Stachys is a of and or . The vary from tall, with simple, , , long with serrate margins. In most species, the leaves are softly hairy. The are long, clustered in the axils of the leaves on the upper part of the stem. The corolla is 5-lobed with the top lobe forming a 'hood', varying from white to pink, purple, red or pale yellow.


Circumscription
The distinction between Stachys and other genera is unclear and has varied from one author to another. In 2002, a molecular phylogenetic showed that Stachys officinalis is not closely related to the rest of the genus.Lindqvist, C. and V. A. Albert. 2002. Origin of the Hawaiian endemic mints within North American Stachys (Lamiaceae). American Journal of Botany 89(10), 1709–24. This study also found six other genera to be within Stachys as it is currently circumscribed. The embedded genera are Prasium, Phlomidoschema, , Haplostachys, , and Stenogyne.


Fossil record
Stachys pliocenica fossil seeds are known from Upper strata of Bulgaria and strata of south-eastern Belarus. The fossil seeds are similar to the seeds of .The Pliocene flora of Kholmech, south-eastern Belarus and its correlation with other Pliocene floras of Europe by Felix Yu. Velichkevich and Ewa Zastawniak - Acta Palaeobot. 43(2): 137–259, 2003


Diversity
Selected species include:


Formerly placed here
  • Agastache foeniculum (Pursh) Kuntze (as S. foeniculum Pursh)
  • Leonurus japonicus Houtt. (as S. artemisia Lour.)
  • (Gouan) P.W.Ball, now in the synonymy of Betonica officinalis
  • Betonica macrantha K.Koch (as S. macrantha (K.Koch) Stearn)
  • Betonica officinalis L. (as S. officinalis (L.) Trevis.)


External links

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