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Horse breeding is reproduction in , and particularly the human-directed process of selective breeding of animals, particularly purebred horses of a given Horse breed. Planned matings can be used to produce specifically desired characteristics in domestication horses. Furthermore, modern breeding management and technologies can increase the rate of conception, a healthy pregnancy, and successful foaling.
A horse is "bred" where it is foaled (born). Thus a colt conceived in England but foaled in the United States is regarded as being bred in the US.Montgomery, E.S, "The Thoroughbred", Arco, New York, 1973 AJC & VRC, "Australian Stud Book", Vol. 31, Ramsay Ware Stockland Pty. Ltd., North Melbourne, 1980 In some cases, most notably in the Thoroughbred breeding industry, American- and Canadian-bred horses may also be described by the state or province in which they are foaled. Some breeds denote the country, or state, where conception took place as the origin of the foal.
Similarly, the "breeder", is the person who owned or leased the mare at the time of foaling. That individual may not have had anything to do with the mating of the mare.Stratton, Charles, The International Horseman's Dictionary, Lansdowne Press, Melbourne, 1978, It is important to review each breed registry's rules to determine which applies to any specific foal.
In the horse breeding industry, the term "half-brother" or "half-sister" only describes horses which have the same dam, but different sires.Summerhayes, RS, Encyclopaedia for Horsemen, Warne & Co, London & New York, 1966 Horses with the same sire but different dams are simply said to be "by the same sire", and no sibling relationship is implied.de Bourg, Ross, "The Australian and New Zealand Thoroughbred", Nelson, West Melbourne, 1980, "Full" (or "own") siblings have both the same dam and the same sire. The terms paternal half-sibling, and maternal half-sibling are also often used. Three-quarter siblings are horses out of the same dam, and are by sires that are either half-brothers (i.e. same dam) or who are by the same sire.Napier, Miles, "Blood Will Tell", JA Allen & Co, London, 1977
and Arabian horse are also classified through the "distaff" or direct female line, known as their "family" or "" line, tracing back to their taproot foundation bloodstock or the beginning of their respective breed registry. The female line of descent always appears at the bottom of a tabulated pedigree and is therefore often known as the bottom line. In addition, the maternal grandfather of a horse has a special term: .
"Linebreeding" technically is the duplication of fourth-generation or more distant ancestors. However, the term is often used more loosely, describing horses with duplication of ancestors closer than the fourth generation. It also is sometimes used as a euphemism for the practice of inbreeding, a practice that is generally frowned upon by horse breeders, though used by some in an attempt to change certain traits.
Once fertilized, the oocyte (egg) remains in the oviduct for approximately 5.5 more days, and then descends into the uterus. The initial single cell combination is already dividing and by the time of entry into the uterus, the egg might have already reached the blastocyst stage.
The gestation period lasts for about eleven months, or about 340 days (normal average range 320–370 days). During the early days of pregnancy, the conceptus is mobile, moving about in the uterus until about day 16 when "fixation" occurs. Shortly after fixation, the embryo proper (so called up to about 35 days) will become visible on trans-rectal ultrasound (about day 21) and a heartbeat should be visible by about day 23. After the formation of the and early placentation is initiated (35–40 days of gestation) the terminology changes, and the embryo is referred to as a fetus. True implantation – invasion into the endometrium of any sort – does not occur until about day 35 of pregnancy with the formation of the endometrial cups, and true placentation (formation of the placenta) is not initiated until about day 40-45 and not completed until about 140 days of pregnancy. The fetus's sex can be determined by day 70 of the gestation using ultrasound. Halfway through gestation the fetus is the size of between a rabbit and a beagle. The most dramatic fetal development occurs in the last 3 months of pregnancy when 60% of fetal growth occurs.
Colts are carried on average about 4 days longer than fillies.
Mares can be used for riding or driving during most of their pregnancy. Exercise is healthy, though should be moderated when a mare is heavily in foal. Exercise in excessively high temperatures has been suggested as being detrimental to pregnancy maintenance during the embryonic period;Mortensen C, Choi YH, Hinrichs K, Ing N, Kraemer D, Vogelsang S, Vogelsang M. 2006. Effects of exercise on embryo recovery rates and embryo quality in the horse. Animal Repro. Sci. 94:395-397 however ambient temperatures encountered during the research were in the region of 100 degrees F and the same results may not be encountered in regions with lower ambient temperatures.
During the first several months of pregnancy, the equine nutrition requirements do not increase significantly since the rate of growth of the fetus is very slow. However, during this time, the mare may be provided supplemental vitamins and minerals, particularly if forage quality is questionable. During the last 3–4 months of gestation, rapid growth of the fetus increases the mare's nutritional requirements. Energy requirements during these last few months, and during the first few months of lactation are similar to those of a horse in full training. Trace minerals such as copper are extremely important, particularly during the tenth month of pregnancy, for proper skeletal formation. Many feeds designed for pregnant and lactation mares provide the careful balance required of increased protein, increased calories through extra fat as well as vitamins and minerals. Overfeeding the pregnant mare, particularly during early gestation, should be avoided, as excess weight may contribute to difficulties foaling or fetal/foal related problems.
On the other hand, some breeders, particularly those in remote areas or with extremely large numbers of horses, may allow mares to foal out in a field amongst a herd, but may also see higher rates of foal and mare mortality in doing so.
Most mares foal at night or early in the morning, and prefer to give birth alone when possible. Labor is rapid, often no more than 30 minutes, and from the time the feet of the foal appear to full delivery is often only about 15 to 20 minutes. Once the foal is born, the mare will lick the newborn foal to clean it and help blood circulation. In a very short time, the foal will attempt to stand and get milk from its mother. A foal should stand and nurse within one hour after birth.
To create a bond with her foal, the mare licks and nuzzles the foal, enabling her to distinguish the foal from others. Some mares are aggressive when protecting their foals, and may attack other horses or unfamiliar humans that come near their newborns.
After birth, a foal's navel is dipped in antiseptic to prevent infection. The foal is sometimes given an enema to help clear the meconium from its digestive tract. The newborn is monitored to ensure that it stands and nurses without difficulty. While most horse births happen without complications, many owners have first aid supplies prepared and a veterinarian on call in case of a birthing emergency. People who supervise foaling should also watch the mare to be sure that she passes the placenta in a timely fashion, and that it is complete with no fragments remaining in the uterus. Retained fetal membranes can cause a serious inflammatory condition (endometritis) and/or infection. If the placenta is not removed from the stall after it is passed, a mare will often eat it, an instinct from the wild, where blood would attract predators.
For example, the Arabian horse of the desert naturally developed speed and endurance to travel long distances and survive in a harsh environment, and domestication by humans added a trainable disposition to the animal's natural abilities. In the meantime, in northern Europe, the locally adapted heavy horse with a thick, warm coat was domesticated and put to work as a farm animal that could pull a plow or wagon. This animal was later adapted through selective breeding to create a strong but rideable animal suitable for the heavily armored knight in war horse.
Then, centuries later, when people in Europe wanted faster horses than could be produced from local horses through simple selective breeding, they imported Arabian horse and other oriental horses to breed as an outcross to the heavier, local animals. This led to the development of breeds such as the Thoroughbred, a horse taller than the Arabian and faster over the distances of a few miles required of a European race horse or light cavalry horse. Another cross between oriental and European horses produced the Andalusian horse, a horse developed in Spain that was powerfully built, but extremely nimble and capable of the quick bursts of speed over short distances necessary for certain types of combat as well as for tasks such as bullfighting.
Later, the people who settled America needed a hardy horse that was capable of working with cattle. Thus, Arabians and Thoroughbreds were crossed on Spanish horses, both domesticated animals descended from those brought over by the Conquistadors, and feral horses such as the Mustangs, descended from the Spanish horse, but adapted by natural selection to the ecology and climate of the west. These crosses ultimately produced new breeds such as the American Quarter Horse and the Criollo of Argentina. In Canada, the Canadian Horse descended from the French stock Louis XIV sent to Canada in the late 17th century.6 The initial shipment, in 1665, consisted of two stallions and twenty mares from the Royal Stables in Normandy and Brittany, the centre of French horse breeding.7 Only 12 of the 20 mares survived the trip. Two more shipments followed, one in 1667 of 14 horses (mostly mares, but with at least one stallion), and one in 1670 of 11 mares and a stallion. The shipments included a mix of draft horses and light horses, the latter of which included both pacing and trotting horses.1 The exact origins of all the horses are unknown, although the shipments probably included Bretons, Normans, Arabians, Andalusians and Barbs.
In modern times, these breeds themselves have since been selectively bred to further specialize at certain tasks. One example of this is the American Quarter Horse. Once a general-purpose working ranch horse, different bloodlines now specialize in different events. For example, larger, heavier animals with a very steady attitude are bred to give competitors an advantage in events such as team roping, where a horse has to start and stop quickly, but also must calmly hold a full-grown steer at the end of a rope. On the other hand, for an event known as cutting, where the horse must separate a cow from a herd and prevent it from rejoining the group, the best horses are smaller, quick, alert, athletic and highly trainable. They must learn quickly, have conformation that allows quick stops and fast, low turns, and the best competitors have a certain amount of independent mental ability to anticipate and counter the movement of a cow, popularly known as "cow sense."
Another example is the Thoroughbred. While most representatives of this breed are bred for horse racing, there are also specialized bloodlines suitable as or show jumping. The hunter must have a tall, smooth build that allows it to trot and canter smoothly and efficiently. Instead of speed, value is placed on appearance and upon giving the equestrian a comfortable ride, with natural jumping ability that shows bascule and good form.
A show jumping, however, is bred less for overall form and more for power over tall fences, along with speed, scope, and agility. This favors a horse with a good galloping stride, powerful hindquarters that can change speed or direction easily, plus a good shoulder angle and length of neck. A jumper has a more powerful build than either the hunter or the racehorse.
One of the earliest people known to document the breedings of their horses were the Bedouin of the Middle East, the breeders of the Arabian horse. While it is difficult to determine how far back the Bedouin passed on pedigree information via an oral tradition, there were written pedigrees of Arabian horses by CE 1330. The Akhal-Teke of West-Central Asia is another breed with roots in ancient times that was also bred specifically for war and racing. The nomads of the steppes bred horses for several thousand years as well, and the Caspian horse is believed to be a very close relative of Ottoman horses from the earliest origins of the Turks in Central Asia.
The types of horse bred varied with culture and with the times. The uses to which a horse was put also determined its qualities, including smooth amblers for riding, fast horses for carrying messengers, heavy horses for plowing and pulling heavy wagons, ponies for hauling cars of ore from mines, packhorses, carriage horses and many others.
Middle Ages Europe bred large horses specifically for war, called . These horses were the ancestors of the great heavy horses of today, and their size was preferred not simply because of the weight of the armor, but also because a large horse provided more power for the knight's lance. Weighing almost twice as much as a normal riding horse, the destrier was a powerful weapon in battle meant to act like a giant battering ram that could quite literally run down men on an enemy line.
On the other hand, during this same time, lighter horses were bred in northern Africa and the Middle East, where a faster, more agile horse was preferred. The lighter horse suited the raids and battles of desert people, allowing them to outmaneuver rather than overpower the enemy. When Middle Eastern warriors and European knights collided in warfare, the heavy knights were frequently outmaneuvered. The Europeans, however, responded by crossing their native breeds with oriental horse such as the Arabian, Barb, and Turkoman horse This cross-breeding led both to a nimbler war horse, such as today's Andalusian horse, but also created a type of horse known as a Courser, a predecessor to the Thoroughbred, which was used as a message horse.
During the Renaissance, horses were bred not only for war, but for haute ecole riding, derived from the most athletic movements required of a war horse, and popular among the elite nobility of the time. Breeds such as the Lipizzan and the now extinct Neapolitan horse were developed from Spanish-bred horses for this purpose, and also became the preferred mounts of cavalry officers, who were derived mostly from the ranks of the nobility. It was during this time that firearms were developed, and so the light cavalry horse, a faster and quicker war horse, was bred for "shoot and run" tactics rather than the shock action as in the Middle Ages. Fine horses usually had a well muscled, curved neck, slender body, and sweeping mane, as the nobility liked to show off their wealth and breeding in paintings of the era.
After Charles II retook the British throne in 1660, horse racing, which had been banned by Cromwell, was revived. The Thoroughbred was developed 40 years later, bred to be the ultimate racehorse, through the lines of three foundation Arabian horse stallions and one Turkish horse.
In the 18th century, James Burnett, Lord Monboddo noted the importance of selecting appropriate parentage to achieve desired outcomes of successive generations. Monboddo worked more broadly in the abstract thought of species relationships and evolution of species. The Thoroughbred breeding hub in Lexington, Kentucky was developed in the late 18th century, and became a mainstay in American racehorse breeding.
The 17th and 18th centuries saw more of a need for fine carriage horses in Europe, bringing in the dawn of the warmblood. The warmblood breeds have been exceptionally good at adapting to changing times, and from their coach horse beginnings they easily transitioned during the 20th century into a sport horse type. Today's warmblood breeds are frequently used in combined driving, but are more often seen competing in show jumping or dressage.
The Thoroughbred continues to dominate the horse racing world, although its lines have been more recently used to improve warmblood breeds and to develop sport horses. The French saddle horse is an excellent example as is the Irish Sport Horse, the latter being an unusual combination between a Thoroughbred and a draft breed.
The American Quarter Horse was developed early in the 18th century, mainly for quarter racing (racing ¼ of a mile). Colonists did not have racetracks or any of the trappings of Europe that the earliest Thoroughbreds had at their disposal, so instead the owners of Quarter Horses would run their horses on roads that lead through town as a form of local entertainment. As the USA expanded West, the breed went with settlers as a farm and ranch animal, and "cow sense" was particularly valued: their use for herding cattle increased on rough, dry terrain that often involved sitting in the saddle for long hours.
However, this did not mean that the original ¼-mile races that colonists held ever went out of fashion, so today there are three types: the stock horse type, the racer, and the more recently evolving sport type. The racing type most resembles the finer-boned ancestors of the first racing Quarter Horses, and the type is still used for ¼-mile races. The stock horse type, used in western events and as a farm and patrol animal is bred for a shorter stride, an ability to stop and turn quickly, and an unflappable attitude that remains calm and focused even in the face of an angry charging steer. The first two are still to this day bred to have a combination of explosive speed that exceeds the Thoroughbred on short distances clocked as high as 55 mph, but they still retain the gentle, calm, and kindly temperament of their ancestors that makes them easily handled.
The Canadian horse's origin corresponds to shipments of French horses, some of which came from Louis XIV's own stable. These were ill-suited to farm work and to the hardscrabble life of the New World, so like the Americans, early Canadians crossed their horses with natives escapees. In time they evolved along similar lines as the Quarter Horse to the South as both the US and Canada spread westward and needed a calm and tractable horse versatile enough to carry the farmer's son to school but still capable of running fast and running hard as a cavalry horse, a stockhorse, or a horse to pull a conestoga wagon.
Other horses from North America retained a hint of their mustang origins by being either derived from stock that Native Americans bred that came in a rainbow of color, like the Appaloosa and American Paint Horse, with those East of the Mississippi River increasingly bred to impress and mimic the trends of the upper classes of Europe: The Tennessee Walking Horse and Saddlebred were originally plantation horses bred for their gait and comfortable ride in the saddle as a plantation master would survey his vast lands like an English lord.
Horses were needed for heavy draft and carriage work until replaced by the automobile, truck, and tractor. After this time, draft and carriage horse numbers dropped significantly, though light riding horses remained popular for recreational pursuits. Draft horses today are used on a few small farms, but today are seen mainly for pulling and plowing competitions rather than farm work. Heavy harness horses are now used as an outcross with lighter breeds, such as the Thoroughbred, to produce the modern warmblood breeds popular in sport horse disciplines, particularly at the Olympic level.
After the mare is bred or artificially inseminated, she is checked using ultrasound 14–16 days later to see if she "took", and is pregnant. A second check is usually performed at 28 days. If the mare is not pregnant, she may be bred again during her next cycle.
It is considered safe to breed a mare to a stallion of much larger size. Because of the mare's type of placenta and its attachment and blood supply, the foal will be limited in its growth within the uterus to the size of the mare's uterus, but will grow to its genetic potential after it is born. Test breedings have been done with draft horse stallions bred to small mares with no increase in the number of difficult births.Stanford, Dr. David, Woodside Equine Clinic, Ashland, VA
When it has been determined that the mare is ready, both the mare and intended stud will be cleaned. The mare will then be presented to the stallion, usually with one handler controlling the mare and one or more handlers in charge of the stallion. Multiple handlers are preferred, as the mare and stallion can be easily separated should there be any trouble.
The Jockey Club, the organization that oversees the Thoroughbred industry in the United States, requires all registered foals to be bred through live cover. Artificial insemination, listed below, is not permitted.Section V, Rule 1, Part D, The American Stud Book Principal Rules and Requirements. The Jockey Club, 2011. Accessed 2011-02-15. Similar rules apply in other countries, such as Australia.See Rule AR.15C, Australian Rules of Racing, which explicitly prohibits human manipulation of the breeding process.
By contrast, the U.S. standardbred industry allows registered foals to be bred by live cover, or by artificial insemination (AI) with fresh or frozen (not dried) semen. No other artificial fertility treatment is allowed. In addition, foals bred via AI of frozen semen may only be registered if the stallion's sperm was collected during his lifetime, and used no later than the calendar year of his death or castration.Rule 26, Section 6, Rules and Regulations of the United States Trotting Association 2009. United States Trotting Association, 2009. Accessed 2011-02-15.
Artificial insemination (AI) has several advantages over live cover, and has a very similar conception rate:
A stallion is usually trained to mount a phantom (or dummy) mare, although a live mare may be used, and he is most commonly collected using an artificial vagina (AV) which is heated to simulate the vagina of the mare. The AV has a filter and collection area at one end to capture the semen, which can then be processed in a lab. The semen may be chilled or frozen and shipped to the mare owner or used to breed mares "on-farm". When the mare is in heat, the person inseminating introduces the semen directly into her uterus using a syringe and pipette.
The world's first cloned horse, Prometea, was born in 2003. Other notable instances of horse cloning are:
Advanced reproductive techniques
Often an owner does not want to take a valuable competition mare out of training to carry a foal. This presents a problem, as the mare will usually be quite old by the time she is retired from her competitive career, at which time it is more difficult to impregnate her. Other times, a mare may have physical problems that prevent or discourage breeding. However, there are now several options for breeding these mares. These options also allow a mare to produce multiple foals each breeding season, instead of the usual one. Therefore, mares may have an even greater value for breeding.
See also
Further reading
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