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The common eland ( Taurotragus oryx), also known as the southern eland or eland antelope, is a large and found in and . An adult male is around tall at the shoulder and can weigh up to with a typical range of . Females are around tall and weigh . It was scientifically described by Peter Simon Pallas in 1766.

Mainly a , its diet is primarily grasses and leaves. Common elands form herds of up to 500 animals, but are not territorial. The common eland prefers with a wide variety of such as savannah, , and open and ; it avoids dense . It uses loud barks, visual and postural movements, and the to communicate and warn others of danger. The common eland is used by humans for , and meat and has been in southern Africa. Eland milk contains more than cow's milk, and can be stored for up to eight months.

It is native to , , the Democratic Republic of the Congo, , , , , , , , , , , , , , and , but is no longer present in . While the common eland's population is decreasing, it is classified as of by the International Union for Conservation of Nature.


Etymology
The scientific name of the common eland is Taurotragus oryx, composed of three words: tauros, tragos, and oryx. Tauros is for a bull or bullock, meaning the same as the taurus. Tragos is Greek for a male goat, referring to the tuft of hair that grows in the eland's ear and its resemblance to a goat's beard. Oryx is Latin and Greek (: orygos) for , referring to the pointed horns of North African antelopes like the common eland and scimitar-horned oryx.

The name "eland" is Afrikaans for "elk" or "moose", from Dutch eland, from obsolete German Elend, probably from obsolete Lithuanian ellenis. When settlers came to the Cape of Good Hope, creating the Dutch Cape Colony, they named the animal after the large, herbivorous . In Dutch, the animal is called "eland antelope" to distinguish it from the moose, which is found in the northern boreal forests.


Physical description
Common elands are spiral-horned antelopes. They are sexually dimorphic, with females being smaller than the males. Females weigh , measure from the snout to the base of the tail and stand at the shoulder. Bulls weigh ,
(1997). 9780691116921, Princeton University Press.
are from the snout to the base of the tail and stand at the shoulder. The tail is long. Male elands can weigh up to .

Their coat differs geographically, with elands in northern part of their range having distinctive markings (torso stripes, markings on legs, dark garters and a spinal crest) that are absent in the south.

(1999). 9780958322331, Chelsea Green Publishing.
Apart from a rough mane, the coat is smooth. Females have a tan coat, while the coats of males are darker, with a bluish-grey tinge. Bulls may also have a series of vertical white stripes on their sides (mainly in parts of the in ). As males age, their coat becomes more grey. Males also have dense on their foreheads and a large on their throats.

Both sexes have horns with a steady spiral ridge (resembling that of the ). The horns are visible as small buds in newborns and grow rapidly during the first seven months. The horns of males are thicker and shorter than those of females (males' horns are long and females' are long), and have a tighter spiral. Males use their horns during rutting season to wrestle and with rivals, while females use their horns to protect their young from predators.

The common eland is the slowest antelope, with a peak speed of that tires them quickly. However, they can maintain a trot indefinitely. Elands are capable of jumping up to from a standing start when startled (up to for young elands). The common eland's life expectancy is generally between 15 and 20 years; in captivity, some live up to 25 years.

Eland herds are accompanied by a loud clicking sound that has been subject to considerable speculation. The weight of the animal may cause the two halves of its to splay apart, and the clicking is the result of the hoof snapping together when the animal raises its leg.

(2026). 9781770092402, Jacana Media.
The sound carries some distance from a herd, and may be a form of communication.


Taxonomy
The common eland was first described in 1766 by German and Peter Simon Pallas. It belongs to the order , family , and subfamily . Common elands are sometimes considered part of the on the basis of molecular phylogenetics, but are usually categorized as , along with the ( T. derbianus).


Subspecies
Three subspecies of common elands have been recognized, though their validity has been disputed.
(2026). 9780521844185, Cambridge University Press.

  • T. o. oryx (Pallas, 1766; Cape eland): also called alces, barbatus, canna and oreas. It is found in and southwestern Africa. The fur is tawny and adults lose their stripes.
  • T. o. livingstonii (, 1864; Livingstone's eland): also called kaufmanni, niediecki, selousi and triangularis. It is found in the Central Zambezian miombo woodlands. Livingstone's eland has brown fur with up to 12 stripes.
  • T. o. pattersonianus (, 1906; East African eland or Patterson's eland): also called billingae. It is found in , hence its common name. Like Livingstone's eland, its fur can also have up to 12 stripes.
Found by and named after John Henry Patterson, who describes the specimen in The Man-eaters of Tsavo (1907).


Genetics and evolution
Male elands have 31 chromosomes and females have 32. The male (Y) chromosome has been translocated to the of an . Both the and late; they do not match well and are variable. The chromosomes resemble those of the ( Tragelaphus strepsiceros).
(2026). 9781421400938, Johns Hopkins University Press.

Male elands and female can produce a viable male hybrid, though whether it is sterile is unknown. An accidental crossing of an East African common eland ( T. o. ') with an East African kudu ( T. s. bea) occurred in the San Diego Zoo Safari Park. This was believed to be due to the absence of male kudus in the herd. The hybrid produced was sterile. The study confirmed the chromosome numbers of both the eland and the kudu and the strangeness of their attached Y chromosomes. Attempt matings of male elands with ( Bos primigenius) and cows ( Bos indicus) indicated the species are not able to interbreed. Female elands have acted as for bongos.

The Bovidae family ancestors of the common eland arose approximately 20 million years ago in Africa; are found throughout Africa and France, but the best records are in sub-Saharan Africa. The first members of the tribe Tragelaphini appear six million years ago during the . An extinct ancestor of the common eland ( Taurotragus arkelli) appears in the in northern and the first T. oryx fossil appears in the in .

In 2010, a genetic study was made basing on the evolutionary history of common elands. Using material from East and Southern Africa, analysis of mitochondrial DNA control-region fragments from 122 individuals revealed information concerning the phylogeography, genetic diversity, and demographic history of the species. The conclusions supported the presence of a long-standing population in the southern Africa and a mosaic of Pleistocene refugia in eastern Africa. The similarity of dates obtained from other studies indicates a significant genetic event around 200 ka.


Habitat and distribution
Common elands live on the open plains of Southern Africa and along the foothills of the great southern African plateau. The species extends north into and most arid zones of , west into eastern and , and south to . However, a low density of elands exists in Africa due to poaching and human settlement.

Elands prefer to live in semiarid areas that contain many shrub-like bushes, and often inhabit grasslands, woodlands, subdesert, bush, and mountaintops with altitudes of about . Elands do, however, avoid forests, swamps and deserts. The places inhabited by elands generally contain , , , , , , and trees and shrubs; some of these also serve as their food.

Eland can be found in many national parks and reserves today, including Nairobi National Park and Tsavo East National Park, Tsavo West National Park, Masai Mara National Reserve, (Kenya); Serengeti National Park, Ruaha National Park and Tarangire National Park, Ngorongoro Crater, (Tanzania); Kagera National Park (Rwanda); Nyika National Park (Malawi); Lake Mburo National Park (Uganda); Kidepo Valley National Park (Uganda); and Kafue National Park (Zambia); Hwange National Park, Matobo National Park, Tuli Safari Area and Chimanimani Eland Sanctuary (Zimbabwe); Kruger National Park, Kgalagadi Transfrontier Park, Giant's Castle and Suikerbosrand NR (South Africa).

They live on home ranges that can be 200–400 km2 for females and juveniles and 50 km2 for males.


Ecology and behavior
Common elands are and . They eat in the morning and evening, rest in shade when hot, and remain in sunlight when cold. They are commonly found in herds numbering up to 500, with individual members remaining in the herd from several hours to several months. Juveniles and mothers tend to form larger herds, while males may separate into smaller groups or wander individually. During , mainly in the rainy season, groups tend to form more regularly. In Southern Africa, common elands will often associate with herds of , and .
(2026). 9780761472667, Marshall Cavendish.

Common elands communicate via gestures, vocalizations, , and display behaviors. The also occurs, primarily in males in response to contact with female urine or genitals. Females urinate to indicate fertility during the appropriate phase of their estrous cycle, as well as to indicate their lack of fertility when harassed by males. If eland bulls find any of their predators nearby, they bark and attempt to attract the attention of others by trotting back and forth until the entire herd is conscious of the danger. Some of their main include , African wild dogs, , and . Eland calves are more vulnerable than adults to their predators.


Diet
Common elands are that browse during drier winter, but have also adapted to during the rainy season when grasses are more common and nutritious. They require a high- diet of leaves from , but will consume lower-quality plant material if available, including , trees, shrubs, grasses, seeds, and tubers. Grasses that elands eat include and and fruits from and . Large antelope can survive on lower-quality food in times of little rain.

Most of their water is obtained from their food, though they drink water when available. As they quickly adjust to the surroundings due to seasonal changes and other causes, they change their feeding habits. They also use their horns to break off branches that are hard to reach.

(2026). 9781615304653, Britannica Educational Pub..


Thermoregulation
Common elands have several thermoregulatory adaptations to help them withstand the extreme temperatures of their environments. Using peripheral thermal receptors on the skin, elands can sense heat and increase or decrease evaporative cooling accordingly. On sunnier days, common elands maintain a cooler skin temperature relative to their inner body temperature. Elands achieve cooler skin temperatures by increasing cutaneous . This allows them to feel cooler, even though their internal body temperature stays relatively the same throughout the day. The eland can also conserve water by increasing its body temperature. When temperatures rise above a certain threshold, an increase in and panting is also observed. Common elands use their sparse fur coats to dissipate excess heat via reradiation. The is also believed to play a role in thermoregulation. Due to its high surface area to volume ratio, it may allow for efficient thermoregulation in larger common elands with larger dewlaps.


Sociability and reproduction
Females are at 15 to 36 months and males at 4 to 5 years. may occur anytime after reaching sexual maturity, but is mostly seen in the rainy season. In Zambia, young are born in July and August, while elsewhere this is the mating season. Mating begins when elands gather to feed on lush, green plains with plentiful grass, and some males and females start mating with each other in separate pairs. Males chase the females to find out if they are in . They also test the female's urine. Usually, a female chooses the most dominant and fit male to mate with. Sometimes, she runs away from males trying to mate, causing more attraction. This results in fights between males, in which their hard horns are used. A female allows a male to mount after two to four hours. Males usually keep close contact with females in the mating period. The dominant male can mate with more than one female. Females have a period of nine months, and give birth to only one calf each time.

Males, females, and juveniles each form separate social groups. The male groups are the smallest; the members stay together and search for food or water sources. The female group is much larger and covers greater areas. They travel the grassy plains in wet periods and prefer bushy areas in dry periods. Females have a complex . The nursery and juvenile group is naturally formed when females give birth to calves. After about 24 hours of the delivery, the mother and calf join this group. The calves start befriending each other and stay back in the nursery group, while the mother returns to the female group. The calves leave the nursery group when they are at least two years old and join a male or female group.


Diseases and parasites
Common elands are resistant to , a protozoan infection that has the as a vector, but not to the -transmitted disease theileriosis. The disease-causing bacterium Theileria taurotragi has caused many eland deaths. Clostridium chauvoei, another bacterium, can be harmful, as well. Elands are also hosts to several kinds of . In one study, an eland was found to be host to the species A. gemma and A. variegatum, and species R. decoloratus, R. appendiculatus, R. evertsi, R. pulchellus and R. pravus. Elands produce for bacteria, but none for Mycobacterium paratuberculosis or various types of like contagious bovine pneumonia and contagious caprine pneumonia, normally infectious in cows or antelopes.


Interaction with humans

Conservation
Currently, common elands are not endangered. They are conserved by the United States Endangered Species Act, and regulated in international trade by the . A 2016 assessment for the International Union for Conservation of Nature (IUCN) estimates the number of mature individuals to be 90,000-110,000, with populations considered stable or increasing in the countries of Namibia, Botswana, Zimbabwe, South Africa, Malawi, Kenya and Tanzania.

The population is, however, gradually decreasing due to habitat loss, caused by expanding human settlements and poaching for its superior meat.

(1990). 9782831700168, IUCN.
As they are docile and inactive most of the time, they can easily be killed. The species became extinct in and , but has been reintroduced.

The IUCN states that about half of the estimated total population lives in protected areas and 30% on private land. Protected areas that support major populations include Omo (Ethiopia), , Katavi, Ruaha, and Selous-Kilombero (Tanzania), and North Luangwa (), (), (), Kgalagadi Transfrontier Park (/) and Ukhahlamba Drakensberg Park (South Africa). Most of these populations appear to be stable. Relatively large numbers of common elands now live on private land, particularly in Namibia, Zimbabwe, and South Africa, reflecting its value as a trophy animal. Common elands have also been widely domesticated in Zimbabwe, South Africa, Kenya, Russia, and Ukraine.

(1999). 9782831704777, The IUCN Species Survival Commission.


Uses
The common eland is sometimes farmed and hunted for its meat, and in some cases can be better used than cattle because it is more suited to African climates. This has led to some Southern African farmers switching from to eland. Common elands are also pictured as supporters in the coat of arms of , Namibia.


Husbandry
Common elands have a mild and have been successfully for meat and milk production in South Africa and Russia. Their need for water is quite low because they produce with a high content, but they require a substantial grazing area, along with and large amounts of supplementary feed such as , , , and , which can be expensive. A female can produce up to of milk per day that is richer in than cow milk. The pleasant-tasting milk has a content of 11-17% and can be stored for up to eight months if properly prepared, versus several days for cow milk.

Housing common elands is difficult due to their ability to jump over fences as high as or simply break through using their substantial mass. Sometimes, wild elands break through enclosures to mix with domesticated ones. Common elands can reproduce in captivity, but calf survival is low and the young may need to be separated from their mothers to ensure health and adequate feeding. requires care because the generally placid animals startle easily and require large amounts of space.

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