Product Code Database
Beavers (genus Castor) are large, semiaquatic rodents of the Northern Hemisphere. There are two existing species: the North American beaver ( Castor canadensis) and the Eurasian beaver ( C. fiber). Beavers are the second-largest living rodents, after Hydrochoerus, weighing up to . They have stout bodies with large heads, long chisel-like incisors, brown or gray fur, hand-like front feet, Webbed foot back feet, and tails that are flat and scaly. The two species differ in skull and tail shape and fur color. Beavers can be found in a number of freshwater habitats, such as rivers, streams, lakes and ponds. They are herbivorous, consuming tree bark, aquatic plants, grasses and sedges.
Beavers build Beaver dam and lodges using tree branches, vegetation, rocks and mud; they chew down trees for building material. Dams restrict water flow, forming ponds, and lodges (usually built in ponds) serve as shelters. Their infrastructure creates used by many other species, and because of their effect on other organisms in the ecosystem, beavers are considered a keystone species. Adult males and females live in Pair bond with their offspring. After their first year, the young help their parents repair dams and lodges; older siblings may also help raise newly born offspring. Beavers hold territories and mark them using scent mounds made of mud, debris, and castoreum—a liquid substance excreted through the beaver's urethra-based castor sacs. Beavers can also recognize their kin by their anal gland secretions and are more likely to tolerate them as neighbors.
Historically, beavers have been hunted for their fur, meat, and castoreum. Castoreum has been used in medicine, perfume, and food flavoring; beaver pelts have been a major driver of the fur trade. Before protections began in the 19th and early 20th centuries, overhunting had nearly exterminated both species. Their populations have since rebounded, and they are listed as species of least concern by the IUCN Red List of mammals. In human culture, the beaver symbolizes industriousness, especially in connection with construction; it is the national animal of Canada.
There are two Extant taxa species: the North American beaver ( Castor canadensis) and the Eurasian beaver ( C. fiber). The Eurasian beaver is slightly longer and has a more lengthened skull, triangular nasal cavity (as opposed to the square ones of the North American species), a lighter fur color, and a narrower tail.
The more basal castorids had several unique features: more complex occlusion between cheek teeth, parallel rows of upper teeth, premolars that were only slightly smaller than molars, the presence of a third set of premolars (P3), a Foramen in the stapes of the inner ear, a smooth palatine bone (with the palatine opening closer to the rear end of the bone), and a longer snout. More derived castorids have less complex occlusion, upper tooth rows that create a V-shape towards the back, larger second premolars compared to molars, absence of a third premolar set and stapes hole, a more grooved palatine (with the opening shifted towards the front), and reduced incisive foramen. Members of the subfamily Palaeocastorinae appeared in late-Oligocene North America. This group consisted primarily of smaller animals with relatively large front legs, a flattened skull, and a reduced tail—all features of a fossorial (burrowing) lifestyle.
In the early Miocene (about 24 mya), castorids Evolution a semiaquatic lifestyle. Members of the subfamily Castoroidinae are considered to be a sister group to modern beavers, and included giants like Castoroides of North America and Trogontherium of Eurasia. Castoroides is estimated to have had a length of and a weight of . Fossils of one genus in Castoroidinae, Dipoides, have been found near piles of chewed wood, though Dipoides appears to have been an inferior woodcutter compared to Castor. Researchers suggest that modern beavers and Castoroidinae shared a bark-eating common ancestor. Dam and lodge-building likely developed from bark-eating, and allowed beavers to survive in the harsh winters of the subarctic. There is no conclusive evidence for this behavior occurring in non- Castor species.
The genus Castor likely originated in Eurasia. The earliest fossil remains appear to be C. neglectus, found in Germany and dated 12–10 mya. Mitochondrial DNA studies place the common ancestor of the two living species at around 8 mya. The ancestors of the North American beaver would have crossed the Bering Land Bridge around 7.5 mya. Castor may have competed with members of Castoroidinae, which led to niche differentiation. The fossil species C. praefiber was likely an ancestor of the Eurasian beaver. C. californicus from the Early Pleistocene of North America was similar to but larger than the extant North American beaver.
'
Beavers have large skulls with powerful chewing muscles. They have four chisel-shaped incisors that continue to grow throughout their lives. The incisors are covered in a thick tooth enamel that is colored orange or reddish-brown by iron compounds. The lower incisors have roots that are almost as long as the entire lower jaw. Beavers have one premolar and three molars on all four sides of the jaws, adding up to 20 teeth. The molars have meandering ridges for grinding woody material. The eyes, ears and nostrils are arranged so that they can remain above water while the rest of the body is submerged. The nostrils and ears have valves that close underwater, while nictitating membranes cover the eyes. To protect the larynx and trachea from water flow, the epiglottis is contained within the nasal cavity instead of the throat. In addition, the back of the tongue can rise and create a waterproof seal. A beaver's lips can close behind the incisors, preventing water from entering their mouths as they cut and bite onto things while submerged.
The beaver's front feet are dexterous, allowing them to grasp and manipulate objects and food, as well as dig. The hind feet are larger and have webbed foot, and the second innermost toe has a "double nail" used for grooming. Beavers can swim at ; only their webbed hind feet are used to swim, while the front feet fold under the chest.
The beaver's distinctive tail has a conical, muscular, hairy base; the remaining two-thirds of the appendage is flat and scaly. The tail has multiple functions: it provides support for the animal when it is upright (such as when chewing down a tree), acts as a rudder when it is swimming, and stores fat for winter. It also has a countercurrent blood vessel system which allows the animal to lose heat in warm temperatures and retain heat in cold temperatures.
The beaver's sex organs are inside the body, and the male's penis has a cartilaginous baculum. They have only one opening, a cloaca, which is used for reproduction, scent-marking, defecation, and urination. The cloaca evolved secondarily, as most mammals have lost this feature, and may reduce the area vulnerable to infection in dirty water. The beaver's intestine is six times longer than its body, and the caecum is double the volume of its stomach. Microorganisms in the caecum allow them to process around 30 percent of the cellulose they eat. A beaver defecates in the water, leaving behind balls of sawdust. Female beavers have four mammary glands; these produce milk with 19 percent fat, a higher fat content than other rodents. Beavers have two pairs of glands: castor sacs, which are part of the urethra, and . The castor sacs secrete castoreum, a liquid substance used mainly for marking territory. Anal glands produce an oily substance which the beaver uses as a waterproof ointment for its coat. The substance plays a role in individual and family recognition. Anal secretions are darker in females than males among Eurasian beavers, while the reverse is true for the North American species. Compared to many other rodents, a beaver's brain has a hypothalamus that is much smaller than the cerebrum; this indicates a relatively advanced brain with higher intelligence. The cerebellum is large, allowing the animal to move within a three-dimensional space (such as underwater) similar to tree-climbing squirrels. The neocortex is devoted mainly to touch and hearing. Touch is more advanced in the lips and hands than the whiskers and tail. Vision in the beaver is relatively poor; the beaver eye cannot see as well underwater as an otter. Beavers have a good sense of smell, which they use for detecting land predators and for inspecting scent marks, food, and other individuals.
Beavers can hold their breath for as long as 15 minutes but typically remain underwater for no more than five or six minutes. Dives typically last less than 30 seconds and are usually no more than deep. When diving, their heart rate decreases to 60 beats per minute, half its normal pace, and blood flow is directed more towards the brain. A beaver's body also has a high tolerance for carbon dioxide. When surfacing, the animal can replace 75 percent of the air in its lungs in one breath, compared to 15 percent for a human.
The Eurasian beaver's range historically included much of Eurasia, but was decimated by hunting by the early 20th century. In Europe, beavers were reduced to fragmented populations, with combined population numbers being estimated at 1,200 individuals for the Rhône of France, the Elbe in Germany, southern Norway, the Neman and Dnieper in Belarus, and the Voronezh river in Russia. The beaver has since recolonized parts of its former range, aided by conservation policies and reintroductions. Beaver populations now range across western, central, and eastern Europe, and western Russia and the Scandinavian Peninsula. Beginning in 2009, beavers have been successfully reintroduced to parts of Great Britain. , the total Eurasian beaver population in Europe was estimated at over one million. Small native populations are also present in Mongolia and northwestern China; their numbers were estimated at 150 and 700, respectively, . Under New Zealand's Hazardous Substances and New Organisms Act 1996, beavers are classed as a "prohibited new organism" preventing them from being introduced into the country.
Beavers typically select flat landscapes with diverse vegetation close to the water. North American beavers prefer trees being or less from the water, but will roam several hundred meters to find more. Beavers have also been recorded in mountainous areas. Dispersing beavers will use certain habitats temporarily before finding their ideal home. These include small streams, temporary swamps, ditches, and backyards. These sites lack important resources, so the animals do not stay there permanently. Beavers have increasingly settled at or near human-made environments, including agricultural areas, suburbs, golf courses, and shopping malls.
Beavers have an herbivorous and a generalist diet. During the spring and summer, they mainly feed on herbaceous plant material such as leaves, roots, herbs, ferns, grasses, sedges, Nymphaea, Brasenia, Scirpus, and Typha. During the fall and winter, they eat more bark and cambium of woody plants; tree and shrub species consumed include aspen, birch, oak, dogwood, willow and alder. There is some disagreement about why beavers select specific woody plants; some research has shown that beavers more frequently select species which are more easily digested, while others suggest beavers principally forage based on stem size. Beavers may cache their food for the winter, piling wood in the deepest part of their pond where it cannot be reached by other browsers. This cache is known as a "raft"; when the top becomes frozen, it creates a "cap". The beaver accesses the raft by swimming under the ice. Many populations of Eurasian beaver do not make rafts, but forage on land during winter.
Beavers usually live up to 10 years. , , and bears may prey upon them. Beavers are protected from predators when in their lodges, and prefer to stay near water. Parasites of the beaver include the bacteria Francisella tularensis, which causes tularemia; the protozoan Giardia duodenalis, which causes giardiasis (beaver fever); and the beaver beetle and mites of the genus Schizocarpus. They have also been recorded to be infected with the rabies virus.
Beavers start building dams when they hear running water, and the sound of a leak in a dam triggers them to repair it. To build a dam, beavers stack up relatively long and thick logs between the banks. Heavy rocks keep them stable, and grass is packed between them. Beavers continue to pile on more material until the dam slopes in a direction facing upstream. Dams can range in height from to and can stretch from to several hundred meters long. Beaver dams are more effective in trapping and slowly leaking water than man-made concrete dams. Lake-dwelling beavers do not need to build dams.
Beavers make two types of lodges: bank lodges and open-water lodges. Bank lodges are burrows dug along the shore and covered in sticks while the more complex freestanding, open-water lodges are built over a platform of piled-up sticks. The lodge is mostly sealed with mud, except for a hole at the top which acts as an air vent. Both types are accessed by underwater entrances. The above-water space inside the lodge is known as the "living chamber", and a "dining area" may exist close to the water entrance. Families routinely clean out old plant material and bring in new material.
North American beavers build more open-water lodges than Eurasian beavers. Beaver lodges built by new settlers are typically small and sloppy. More experienced families can build structures with a height of and an above-water diameter of . A lodge sturdy enough to withstand the coming winter can be finished in just two nights. Both lodge types can be present at a beaver site. During the summer, beavers tend to use bank lodges to keep cool and use open-water lodges during the winter. The air vent provides ventilation, and newly added carbon dioxide can be cleared in an hour. The lodge remains consistent in oxygen and carbon dioxide levels from season to season.
Beavers in some areas will dig canals connected to their ponds. The canals fill with groundwater and give beavers access and easier transport of resources, as well as allow them to escape predators. These canals can stretch up to wide, deep, and over long. It has been hypothesized that beavers' canals are not only transportation routes but an extension of their "central place" around the lodge and/or food cache. As they drag wood across the land, beavers leave behind trails or "slides", which they reuse when moving new material.
Beaver ponds, and the wetlands that succeed them, remove sediments and pollutants from waterways, and can stop the loss of important soils. These ponds can increase the productivity of freshwater ecosystems by accumulating nitrogen in sediments. Beaver activity can affect the temperature of the water; in northern latitudes, ice thaws earlier in the warmer beaver-dammed waters. Beavers may contribute to climate change. In Arctic areas, the floods they create can cause permafrost to thaw, releasing methane into the atmosphere.
As wetlands are formed and riparian habitats are enlarged, aquatic plants colonize the newly available watery habitat. One study in the Adirondacks found that beaver engineering lead to an increase of more than 33 percent in herbaceous plant diversity along the water's edge. Another study in semiarid eastern Oregon found that the width of riparian vegetation on stream banks increased several-fold as beaver dams watered previously dry terraces adjacent to the stream. Riparian ecosystems in arid areas appear to sustain more plant life when beaver dams are present. Beaver ponds act as a refuge for riverbank plants during wildfires, and provide them with enough moisture to resist such fires. Introduced beavers at Tierra del Fuego have been responsible for destroying the indigenous forest. Unlike trees in North America, many trees in South America cannot grow back after being cut down. Beaver activity impacts communities of aquatic invertebrates. Damming typically leads to an increase of lentic ecosystem species, like dragonflies, oligochaetes, snails, and mussels. This is to the detriment of Lotic ecosystems species like black flies, stoneflies, and Hydropsychidae. Beaver floodings create more dead trees, providing more habitat for terrestrial invertebrates like Drosophila flies and bark beetles, which live and breed in dead wood. The presence of beavers can increase wild salmon and trout populations, and the average size of these fishes. These species use beaver habitats for spawning, overwintering, feeding, and as havens from changes in water flow. The positive effects of beaver dams on fish appear to outweigh the negative effects, such as blocking of migration. Beaver ponds have been shown to be beneficial to frog populations by protecting areas for larvae to mature in warm water. The stable waters of beaver ponds also provide ideal habitat for freshwater turtles.
Beavers help waterfowl by creating increased areas of water. The widening of the riparian zone associated with beaver dams has been shown to increase the abundance and diversity of birds favoring the water's edge, an impact that may be especially important in semi-arid climates. Fish-eating birds use beaver ponds for foraging, and in some areas, certain species appear more frequently at sites where beavers were active than at sites with no beaver activity. In a study of Wyoming streams and rivers, watercourses with beavers had 75 times as many ducks as those without. As trees are drowned by rising beaver impoundments, they become an ideal habitat for woodpeckers, which carve cavities that may be later used by other bird species. Beaver-caused ice thawing in northern latitudes allows Canada geese to nest earlier.
Other semi-aquatic mammals, such as water voles, muskrats, minks, and otters, will shelter in beaver lodges. Beaver modifications to streams in Poland create habitats favorable to bat species that forage at the water surface and "prefer moderate vegetation clutter". Large herbivores, such as some deer species, benefit from beaver activity as they can access vegetation from fallen trees and ponds.
Adult beavers mate with their partners, though partner replacement appears to be common. A beaver that loses its partner will wait for another one to come by. Estrus cycles begin in late December and peak in mid-January. Females may have two to four estrus cycles per season, each lasting 12–24 hours. The pair typically mate in the water and to a lesser extent in the lodge, for half a minute to three minutes.
Up to four young, or kits, are born in spring and summer, after a three or four-month gestation. Newborn beavers are precocial with a full fur coat, and can open their eyes within days of birth. Their mother is the primary caretaker, while their father maintains the territory. Older siblings from a previous litter also play a role.
After they are born, the kits spend their first one to two months in the lodge. Kits suckle for as long as three months, but can eat solid food within their second week and rely on their parents and older siblings to bring it to them. Eventually, beaver kits explore outside the lodge and forage on their own, but may follow an older relative and hold onto their backs. After their first year, young beavers help their families with construction. Beavers sexually mature around 1.5–3 years. They become independent at two years old, but remain with their parents for an extra year or more during times of food shortage, high population density, or drought.
Beavers mark their territories by constructing scent mounds made of mud and vegetation, scented with castoreum. Those with many territorial neighbors create more scent mounds. Scent marking increases in spring, during the dispersal of yearlings, to deter interlopers. Beavers are generally intolerant of intruders and fights may result in deep bites to the sides, rump, and tail. They exhibit a behavior known as the "dear enemy effect"; a territory-holder will investigate and become familiar with the scents of its neighbors and react more aggressively to the scents of strangers passing by. Beavers are also more tolerant of individuals that are their kin. They recognize them by using their keen sense of smell to detect differences in the composition of anal gland secretions. Anal gland secretion profiles are more similar among relatives than unrelated individuals.
Flow devices, like beaver pipes, are used to manage beaver flooding, while fencing and hardware cloth protect trees and shrubs from beaver damage. If necessary, hand tools, heavy equipment, or explosives are used to remove dams. Hunting, trapping, and relocation may be permitted as forms of population control and for removal of individuals. The governments of Argentina and Chile have authorized the trapping of invasive beavers in hopes of eliminating them. The ecological importance of beavers has led to cities like Seattle designing their parks and green spaces to accommodate the animals. The Martinez beavers became famous in the mid-2000s for their role in improving the ecosystem of Alhambra Creek in Martinez, California.
Zoos have displayed beavers since at least the 19th century, though not commonly. In captivity, beavers have been used for entertainment, fur harvesting, and for reintroduction into the wild. Captive beavers require access to water, substrate for digging, and artificial shelters. Grey Owl pioneered beaver conservation in the early 20th century. Belaney wrote several books, and was first to professionally film beavers in their environment. In 1931, he moved to a log cabin in Prince Albert National Park, where he was the "caretaker of park animals" and raised a beaver pair and their four offspring.
Castoreum was used for a variety of medical purposes; Pliny the Elder promoted it as a treatment for stomach problems, flatulence, seizures, sciatica, vertigo, and epilepsy. He stated it could stop hiccups when mixed with vinegar, toothaches if mixed with oil (by administering into the ear opening on the same side as the tooth), and could be used as an antivenom. The substance has traditionally been prescribed to treat hysteria in women, which was believed to have been caused by a "toxic" womb. Castoreum's properties have been credited to the accumulation of salicylic acid from willow and aspen trees in the beaver's diet, and has a physiological effect comparable to aspirin. Today, the medical use of castoreum has declined and is limited mainly to homeopathy. The substance is also used as an ingredient in perfumes and , and as a flavouring in food and drinks.
Various Native American groups have historically hunted beavers for food, they preferred its meat more than other red meats because of its higher calorie and fat content, and the animals remained plump in winter when they were most hunted. The bones were used to make tools. In medieval Europe, the Catholic Church considered the beaver to be part mammal and part fish, and allowed followers to eat the scaly, fishlike tail on meatless Fridays during Lent. Beaver tails were thus highly prized in Europe; they were described by French naturalist Pierre Belon as tasting like a "nicely dressed eel".
Beaver pelts were used to make Beaver hat; felting would remove the guard hairs. The number of pelts needed depended on the type of hat, with Cavalier hat and Capotain hats requiring more fur than . In the late 16th century, Europeans began to deal in North American furs due to the lack of taxes or tariffs on the continent and the decline of fur-bearers at home. Beaver pelts caused or contributed to the Beaver Wars, King William's War, and the French and Indian War; the trade made John Jacob Astor and the owners of the North West Company very wealthy. For Europeans in North America, the fur trade was a driver of the exploration and westward exploration on the continent and contact with native peoples, who traded with them.
The fur trade peaked between 1860 and 1870, when over 150,000 beaver pelts were purchased annually by the Hudson's Bay Company and fur companies in the United States. The contemporary global fur trade is not as profitable due to conservation, anti-fur and animal rights campaigns.
Native American myths emphasize the beaver's skill and industriousness. In the mythology of the Haida people, beavers are descended from the Beaver-Woman, who built a dam on a stream next to their cabin while her husband was out hunting and gave birth to the first beavers. In a Cree story, the Great Beaver and its dam caused a Flood myth. Other tales involve beavers using their tree chewing skills against an enemy. Beavers have been featured as companions in some stories, including a Lakota people tale where a young woman flees from her evil husband with the aid of her pet beaver.
Europeans have traditionally thought of beavers as fantastical animals due to their amphibious nature. They depicted them with exaggerated tusk-like teeth, dog- or pig-like bodies, fish tails, and visible testicles. French cartographer Nicolas de Fer illustrated beavers building a dam at Niagara Falls, fantastically depicting them like human builders. Beavers have also appeared in literature such as Dante Alighieri Divine Comedy and the writings of Athanasius Kircher, who wrote that on Noah's Ark the beavers were housed near a water-filled tub that was also used by and otters.
The beaver has long been associated with Canada, appearing on the first pictorial postage stamp issued in the Canadian colonies in 1851 as the so-called "Three-Penny Beaver". It was declared the national animal in 1975. The five-cent coin, the coat of arms of the Hudson's Bay Company, and the logos for Parks Canada and Roots Canada use its image. Frank and Gordon are two fictional beavers that appeared in Bell Canada advertisements between 2005 and 2008. However, the beaver's status as a rodent has made it controversial, and it was not chosen to be on the Arms of Canada in 1921. The beaver has commonly been used to represent Canada in political cartoons, typically to signify it as a friendly but relatively weak nation. In the United States, the beaver is the state animal of New York and Oregon. It is also featured on the coat of arms of the London School of Economics.
|
|
