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Wrens are a family, Troglodytidae, of small brown birds. The family includes 96 and is divided into 19 . All species are restricted to the except for the that is widely distributed in the . In , the Eurasian wren is commonly known simply as the "wren", as it is the originator of the name. The name wren has been applied to other, unrelated birds, particularly the New Zealand wrens () and the Australian wrens ().

Most wrens are visually inconspicuous though they have loud and often complex songs. Exceptions include the relatively large members of the genus , which can be quite bold in their behaviour. Wrens have short wings that are barred in most species, and they often hold their tails upright. Wrens are primarily , eating insects, spiders and other small , but many species also eat vegetable matter and some eat small frogs and lizards.


Etymology and usage
The English name "wren" derives from and , attested (as wernnaa) very early, in an eighth-century gloss. It is to , wrendilo, and (the latter two including an additional diminutive -ilan suffix). The Icelandic name is attested in () as rindilþvari. This points to a name wrandjan-, but the further etymology of the name is unknown.Kluge-Lutz, English Etymology tentatively suggest association with Old High German (w)renno "stallion", but Suolahti (1909) rejects this as unlikely.

The wren was also known as the kuningilin ('kinglet') in Old High German, a name associated with the of the election of the "king of birds". The bird that could fly to the highest altitude would be made king. The outflew all other birds, but he was beaten by a small bird that had hidden in his plumage. This fable was already known to ( Historia Animalium 9.11)"It goes by the nickname of 'old man' or 'king'; and the story goes that for this reason the eagle is at war with him." http://etext.virginia.edu/etcbin/toccer-new2?id=AriHian.xml&images=images/modeng&data=/texts/english/modeng/parsed&tag=public&part=9&division=div2 and Pliny ( Natural History 10.95),"The roiall Ægle hateth the Wren, and why? because (if we may beleeve it) he is named Regulus, i." http://penelope.uchicago.edu/holland/pliny10.html and was taken up by authors such as Johann Geiler von Kaisersberg, but it most likely originally concerned (Regulus, such as the ) and was apparently motivated by the yellow "crown" sported by these birds (a point noted already by ). The confusion stemmed in part from the similarity and consequent interchangeability of the words for the wren (βασιλεύς basileus, 'king'). and the crest (βασιλίσκος basiliskos, 'kinglet'),.

(2025). 9780415238519, Routledge.
and the legend's reference to the "smallest of birds" becoming king likely led the title to be transferred to the equally tiny wren.
(2025). 9780701169077, Chatto & Windus.
In modern , the name of the bird is Zaunkönig ('king of the fence (or hedge)') and in , the name is winterkoning ('king of winter').

The family name Troglodytidae is derived from troglodyte, which means 'cave-dweller'. Wrens get their scientific name from the tendency of some species to forage in dark crevices. Dated 1807 on title page but not published until 1809.

The name "wren" is also ascribed to other families of passerine birds throughout the world. In Europe, kinglets are occasionally known as "wrens", with the and known as the "fire-crested wren" and "golden-crested wren", respectively.

The 27 "wren" species in the family are unrelated, as are the New Zealand wrens in the family Acanthisittidae, the in the family Thamnophilidae, and the Old World babblers of the family Timaliidae.


Description
Wrens are medium-small to very small birds. The Eurasian wren is among the smallest birds in its range, while the smaller species from the are among the smallest in that part of the world. They range in size from the white-bellied wren, which averages under and , to the , which averages about and weighs almost . The dominating colors of their are generally drab, composed of gray, brown, black, and white, and most species show some barring, especially on the tail or wings. No sexual dimorphism is seen in the plumage of wrens, and little difference exists between young birds and adults. All have fairly long, straight to marginally decurved (downward-curving) bills.

Wrens have loud and often complex songs, sometimes given in by a pair. The songs of members of the genera and have been considered especially pleasant to the human ear, leading to common names such as , , , and southern nightingale-wren.


Distribution and habitat
Wrens are principally a New World family, distributed from and to southern , with the greatest in the . As suggested by its name, the is the only species of wren found outside the Americas, as restricted to Europe, Asia, and northern Africa (it was formerly considered with the and of North America). The insular species include the Clarión wren and from the Revillagigedo Islands in the Pacific Ocean, and Cobb's wren in the , but few islands have a species of wren, with only the southern house wren in the , the of , and the highly restricted in a single swamp in .

The various species occur in a wide range of habitats, ranging from dry, sparsely wooded country to rainforests. Most species are mainly found at low levels, but members of the genus are frequently found higher, and the two members of are restricted to the . A few species, notably the Eurasian wren and the house wren, are often associated with humans. Most species are resident, remaining in Central and South America all year round, but the few species found in temperate regions of the Northern Hemisphere are partially migratory, spending the winter further south.


Behavior and ecology
Wrens vary from highly secretive species such as those found in the genus to the highly conspicuous genus , the members of which frequently sing from exposed perches. The family as a whole exhibits a great deal of variation in their behavior. Temperate species generally occur in pairs, but some tropical species may occur in parties of up to 20 birds.

Wrens build dome-shaped nests, and may be either monogamous or polygamous, depending on species.

(1991). 9781853911866, Merehurst Press.

Though little is known about the feeding habits of many of the Neotropical species, wrens are considered primarily , eating insects, spiders, and other small . Many species also take vegetable matter such as seeds and berries, and some (primarily the larger species) take small frogs and lizards. The Eurasian wren has been recorded wading into shallow water to catch small fish and tadpoles; Sumichrast's wren and the take snails; and the and have been recorded attacking and eating bird eggs (in the latter species, even eggs of conspecifics). A local name for the giant wren and is chupahuevo ('egg-sucker'), but whether the latter actually eats eggs is unclear. The and northern house wren sometimes destroy bird eggs, and the rufous-and-white wren has been recorded killing nestlings, but this is apparently to eliminate potential food competitors rather than to feed on the eggs or nestlings. Several species of Neotropical wrens sometimes participate in mixed-species flocks or , and the Eurasian wren may follow to catch prey items disturbed by them.


Taxonomy and systematics
Revised following Martínez Gómez et al. (2005) and Mann et al. (2006), the taxonomy of some groups is highly complex, and future species-level splits are likely. Additionally, undescribed are known to exist. The black-capped donacobius is an enigmatic species traditionally placed with the wrens more for lack of a more apparent alternative than as a result of thorough study. It was recently determined to be most likely closer to certain , possibly the newly established , and might constitute a family.

The genus level of the Troglodytidae shown below is based on a molecular phylogenetic study by Tyler Imfeld and collaborators that was published in 2024. The number of species in each genus is based on the list maintained by Frank Gill, Pamela C. Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC).

Family Troglodytidae

[[File:Odontorchilus branickii.jpg|thumb|right|200px|[[Grey-mantled wren]] (''Odontorchilus branickii'')]]
     
  • Genus
    • Grey-mantled wren ( Odontorchilus branickii)
    • Tooth-billed wren ( Odontorchilus cinereus)
  • Genus
  • Genus
  • Genus
    • Sumichrast's wren ( Hylorchilus sumichrasti)
    • Nava's wren ( Hylorchilus navai)
  • Genus
  • Genus
    • ( Cistothorus stellaris)
    • Mérida wren or paramo wren ( Cistothorus meridae)
    • Apolinar's wren ( Cistothorus apolinari)
    • ( Cistothorus platensis)
    • ( Cistothorus palustris)
  • Genus
    • Bewick's wren ( Thryomanes bewickii)
  • Genus
  • Genus (formerly included in Thryothorus)
    • Black-throated wren ( Pheugopedius atrogularis)
    • Sooty-headed wren ( Pheugopedius spadix)
    • Black-bellied wren ( Pheugopedius fasciatoventris)
    • Plain-tailed wren ( Pheugopedius euophrys)
    • ( Pheugopedius schulenbergi)
    • ( Pheugopedius eisenmanni)
    • ( Pheugopedius genibarbis)
    • ( Pheugopedius mystacalis)
    • ( Pheugopedius coraya)
    • ( Pheugopedius felix)
    • Spot-breasted wren ( Pheugopedius maculipectus)
    • Rufous-breasted wren ( Pheugopedius rutilus)
    • Speckle-breasted wren ( Pheugopedius sclateri)
  • Genus (formerly included in Thryothorus)
    • ( Thryophilus pleurostictus)
    • Rufous-and-white wren ( Thryophilus rufalbus)
    • ( Thryophilus sernai)
    • Niceforo's wren ( Thryophilus nicefori)
    • ( Thryophilus sinaloa)
  • Genus (formerly included in Thryothorus)
    • Cabanis's wren ( Cantorchilus modestus)
    • ( Cantorchilus zeledoni)
    • ( Cantorchilus elutus)
    • Buff-breasted wren ( Cantorchilus leucotis) (probably not )
    • Superciliated wren ( Cantorchilus superciliaris)
    • Fawn-breasted wren ( Cantorchilus guarayanus)
    • ( Cantorchilus longirostris)
    • ( Cantorchilus griseus)
    • ( Cantorchilus semibadius)
    • ( Cantorchilus nigricapillus)
    • Stripe-breasted wren ( Cantorchilus thoracicus)
    • Stripe-throated wren ( Cantorchilus leucopogon)
  • Genus
    • ( Thryothorus ludovicianus)
      • White-browed wren ( Thryothorus (ludovicianus) albinucha)
  • Genus Troglodytes (10–15 species, depending on taxonomy; includes species sometimes considered to be in the genus Nannus, which may be distinct)
    • ( Troglodytes troglodytes)
    • ( Troglodytes hiemalis)
    • ( Troglodytes pacificus)
    • Clarión wren ( Troglodytes tanneri)
    • House wren ( Troglodytes aedon)
    • Cobb's wren ( Troglodytes cobbi)
    • ( Troglodytes sissonii)
    • Rufous-browed wren ( Troglodytes rufociliatus)
    • ( Troglodytes ochraceus)
    • ( Troglodytes solstitialis)
    • Santa Marta wren ( Troglodytes monticola)
    • ( Troglodytes rufulus)
  • Genus
  • Genus
    • White-bellied wren ( Uropsila leucogastra)
  • Genus (wood wrens)
    • White-breasted wood wren ( Henicorhina leucosticta)
    • Grey-breasted wood wren ( Henicorhina leucophrys)
    • Hermit wood wren ( Henicorhina anachoreta) – split from H. leucophrys
    • Bar-winged wood wren ( Henicorhina leucoptera)
    • Munchique wood wren ( Henicorhina negreti)
  • Genus
    • Northern nightingale-wren ( Microcerculus philomela)
    • Southern nightingale-wren ( Microcerculus marginatus)
    • ( Microcerculus ustulatus)
    • ( Microcerculus bambla)
  • Genus
    • Chestnut-breasted wren ( Cyphorhinus thoracicus)
    • ( Cyphorhinus arada)
    • ( Cyphorhinus phaeocephalus)


Relationship with humans
The wren features prominently in culture. The Eurasian wren has been long considered "the king of birds" in Europe. Killing one or harassing its nest is associated with bad luck, such as broken bones, lightning strikes on homes, or injury to cattle. , celebrated in parts of Ireland on Saint Stephen's Day (26 December), features a fake wren being paraded around town on a decorative pole; up to the 20th century, real birds were hunted for this purpose.
(1997). 9780870499609, University of Tennessee Press.
A possible origin for the tradition is revenge for the betrayal of by a noisy wren when he was trying to hide from enemies in a bush.

The ( Thryothorus ludovicianus) has been the state bird of since 1948, and features on the back of its state quarter. The British farthing featured a wren on the reverse side from 1937 until its demonetisation in 1960. The ( Campylorhynchus brunneicapillus) was designated the state bird of in 1931.

The Women's Royal Naval Service (WRNS) were nicknamed Wrens based on the acronym WRNS. After the Women's Royal Navy Service was integrated into the Royal Navy in 1993, the title of Wren was dropped from official usage, however unofficially female sailors are still referred to as Wrens.

File:2000 SC Proof.png|alt=South Carolina state quarter File:British farthing 1951 reverse.png|alt=British farthing

  • English


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