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   » » Wiki: Turbinicarpus
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Turbinicarpus is a of very small to medium-sized , which inhabit the north-eastern regions of , in particular the states of San Luis Potosí, , Nuevo León, Querétaro, Hidalgo, , and .


Taxonomy
The taxon was first proposed by as Strombocactus subgenus Turbinicarpus. It was elevated to a genus in 1937 by and Backeberg. The circumscription of Turbinicarpus has been described as "remarkably unstable", with species regularly transferred to other genera. Its taxonomic history is often mixed with that of other genera like , , Gymnocactus, , , Normanbokea, , , , and , as the results of almost two centuries of constant evolution in the understanding of the affinities and relationships inside the family Cactaceae. A genus revision by Davide Donati in 2003, and again in 2004 with Carlo Zanovello, was based on a wide range of characters. At the end of that study, was considered a distinct genus from Turbinicarpus. The genus Turbinicarpus was subdivided in two subgenera at the light of the results of the DNA analysis, and into many series because of the ontogeny of the spination. The broad circumscription of Turbinicarpus was recognized as by Hunt in 2016. A study published in 2019 showed that both and Rapicactus were distinct from a more narrowly circumscribed and so monophyletic Turbinicarpus:


Species
, Plants of the World Online accepted the following species and hybrids:

Mexico.
Mexico (Nuevo León)
Mexico (San Luis Potosí)
Mexico (Nuevo León)
Mexico (Hidalgo)
Mexico (Nuevo León)
Mexico (San Luis Potosí)
Mexico (Ciudad del Maiz on the north, and Cerritos-Villa Juarez on the west and Rio Verde )
Mexico Northeast
Mexico(Coahuila, Nuevo León, San Luis Potosí, and Tamaulipas )
Mexico ( San Luis Potosí and Tamaulipas )
Mexico(Nuevo León, San Luis Potosí, Tamaulipas )
Mexico (Rayones, Nuevo León)
Mexico (Coahuila and San Luis Potosí)
Mexico (Tamaulipas, Nuevo León and San Luis Potosí)


Natural hybrids
Mexico (Cerritos, San Lois Potosi )
Mexico (Nuevo León)
Mexico Northeast


Synonymy
The following genera haven been brought into synonymy with Turbinicarpus:
  • Gymnocactus Backeb.
  • Normanbokea Kladiwa & Buxb.


Ecology
These succulent plants grow mostly on limestone soil (never on volcanic soil), at altitudes between 300 and 3300 metres above sea level.

Turbinicarpus species are usually confined to specific habitats, generally hostile for the majority of plants, mostly in very drained rocky areas, composed of limestone, sandstone, schist (neutral or alkaline), or in very acidic and humiferous understorey, or in gypsum veins, sometimes so pure that they are almost white.

In particular, Turbinicarpus sensu stricto is adapted to extreme niches: more than 80% of the species grow in rock cracks or among the pebbles beneath them, where enough dust has accumulated to enable root development. It would seem almost impossible that plants so small could survive in such an environment, however in those species that inhabit dry and exposed areas, the root is very thick, becoming a taproot and acting like an anchor on the slopes but, more important, as water storage for the dry periods, capable of significantly retracting into the ground so that the stem is less exposed to the sun; the spines are often changed to adopt a very papery structure, capable of absorbing good quantities of water. Furthermore, the general look is extremely mimetic, thanks to the epidermis colour and the interlacing spines, guaranteeing a certain protection from eventual herbivores.

  • Edward F. Anderson, The Cactus Family (Timber Press, 2001), pp. 665–673
  • Grupo San Luis, The Genus Turbinicarpus in San Luis Potosì (Ed. Cactus&Co, 2004)


External links

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