Percrocutidae is an extinct family of hyena-like feliform Carnivora endemic to Asia, Africa, and Southern Europe from the Middle Miocene through the Pliocene, existing for about .[ Paleobiology Database: Percrocutidae basic information]
The first percrocutids are known from the middle Miocene of Europe and western Asia and belonged to the genus Percrocuta. Percrocuta already had large , but did not carry such a massive bite as the later form Dinocrocuta, from the later Miocene. Originally, these carnivores were placed with the hyenas in the family Hyaenidae. , most scientists considered the Percrocutidae to be a distinct family that evolved their morphology similar to hyenas due convergent evolution, - although they are usually placed sister-taxa/immediate outgroup to Hyaenidae. Sometimes it was placed with the family Stenoplesictidae into the superfamily Stenoplesictoidea. A 2022 study placed Dinocrocuta and Percrocuta as true hyaenids, which if correct would invalidate the family Percrocutidae.
Taxonomy and evolution
Taxonomic history
Percrocuta was first considered as a side-branch outside of Hyaenidae by Thenius in 1966.
It was later named as a different subfamily, Percrocutinae, of Hyaenidae in 1976, and at that time was proposed to include
Percrocuta,
Adcrocuta eximia, and
Allohyaena kadici.
Dinocrocuta was elevated from a subgenus to a full genus in 1988.
The family Percrocutidae was formally elevated in 1991, to include the genera Percrocuta, Dinocrocuta, Belbus and Allohyaena.
Later studies have suggested that Belbus and Allohyaena are true hyaenids and not percrocutids.
Classification
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| † Percrocuta |
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† P. abessalomi
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† P. carnifex
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† P. grandis
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† P. leakeyi
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† P. miocenica
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† P. tobieni
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† P. tungurensis
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† P. xixiaensis
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The list follows McKenna and Bell's Classification of Mammals for prehistoric genera (1997).[Malcolm C. McKenna, Susan K. Bell: in Columbia University Press, New York 1997, 631 Seiten, ] In contrast to McKenna and Bell's classification, they are not included as a subfamily into the Hyaenidae but as a separate family Percrocutidae.
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Jordi Agustí: Mammoths, Sabertooths and Hominids 65 Million Years of Mammalian Evolution in Europe, Columbia University Press, 2002.