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Megalictis
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Megalictis (meaning "great weasel") is an of large predatory that existed in during the "" from the Late (Ar4) in the epoch. It is thought to have resembled a huge, jaguar-sized , weighing up to .


History of discovery
The genus Megalictis was first described by W. D. Matthew in 1907, and assigned to the family .Paleobiology Database Two similar genera discovered at the same time, Aelurocyon (Peterson, 1907) and Paroligobunis (Peterson, 1910) were identified as synonymous with Megalictis in 1996Andersson, p.39 though Paroligobunis was re-established as a separate genus in 1998. P. R. Bjork, in 1970, assigned the genus to the subfamily ,Bjork, p.18 whilst J. A. Baskin reassigned it to in 1998. Three species have been identified in the genus: M. ferox, M. petersoni, and M. frazieri, whilst two more, Megalictis brevifacies and Megalictis simplicidens, have since been determined to be synonymous with M. ferox. Other synonyms of M. ferox include Aelurocyon brevifacies, Brachypsalis simplicidens and Paroligobunis simplicidens.

Specimens of M. ferox have been found at ten sites in , and , dated to the Harrisonian period - from 24.8 to 20.6 million years ago. The 19 specimens so far identified have a wide range of sizes, which is believed to be evidence of extreme sexual dimorphism, a phenomenon where males and females of the species will grow to different sizes; a 1996 study rejected the possibility of multiple "morphologically uniform" but differently sized species as unlikely.Hunt & Skolnick, quoted Andersson, p.39-40 However, later studies identified significant morphological variation in the joints of the forearms, which would not be consistent with sexual dimorphism. Andersson suggests that the identification of all of these as a single species may need revision.Andersson, p.11 & p.40


Description
Megalictis was one of the largest terrestrial mustelids ever recorded, surpassing in size other mustelids like , , and . Recovered skulls of Megalictis ferox had similar condylobasal lengths to small modern jaguars, ranging from in length, and were almost as wide in mastoid width as those of some , ranging from wide. The enlarged anterior edge of the cornoid process confers a wide gape relative to its short rostral length, suggesting Megalictis possessed a powerful bite force; together with blunt, stout postcanine teeth, Megalictis was well equipped to engage in , similar to modern and . It has been described as having short stout legs, with short feet and long non-retractible claws, similar to a modern but about twice the size, and noted that "the teeth indicate an animal fully as predaceous as the wolverine, but the skeleton points to more fossorial habits".Quoted in Gratacap, p.921 Research comparing the joints of the forelegs to modern mammalian carnivores suggests that it waited in ambush and grappled with its prey, rather than hunting it down by pursuit like a modern .Hopkin


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