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Hexacorallia (synonym: Zoantharia

(2016). 9781605353753, Sinauer.
(2025). 9780030259821, Cengage Learning.
[1]
Grzimek, Bernhard (2003). Grzimek's Animal Life Encyclopedia: Lower metazoans and lesser deuterosomes. Gale. p. 103. ISBN 978-0-7876-7750-3.PARKER, S. P. (ed.) Synopsis and Classification of Living Organisms 1. New York etc: McGraw-Hill Book Company, 1982. p. 704; older synonyms: Hexacoralla,ZORKOVSKÝ, B. Historická geológia. Bratislava: SVTL, 1958, p. 55 Hexaradiata,HAEFELFINGER, Hans Rudolf, Die Hohltiere. In: Eibl-Eibesfeldt, I., Grzimek, B. et al. (eds.) Grzimeks Tierleben, Band 1 Niedere Tiere. Zürich: Kindler. 1971. pp. 176-272, 521 et seqq. Hexactinia, ActinanthidaDELAGE, Yves, HÉROUARD, Edgard. Traité de zoologie concrète. Tome II, 2eme partie. Paris, 1901 [2]; in English: hexacorals,
(2005). 9780080460123, Elsevier. .
hexacorallians, zoantharians) is a class of .


Other meanings
Both names, i. e. Hexacorallia and Zoantharia, and their English equivalents, also have narrower meanings - see below in the chapter Taxonomy. The name Hexacorallia also has a rare meaning unrelated to the content of this article - it is used as a synonym of the name .


Characteristics
Hexacorals are aquatic organisms formed of polyps, generally with 6-fold symmetry. The number of hexacoral species is approximately 4,300 (as estimated in 1987). Extant hexacorals are composed of 6 orders: the (Ceriantharia), (Zoanthiniaria), (Actiniaria), (Corallimorpharia), the (Antipatharia) and the (Scleractinia). The first four orders are skeleton-less, while the last two orders have a skeleton. Some of the Scleractinia are builders.

The Hexacorallia are distinguished from another class of Anthozoa, , in having six or fewer axes of symmetry in their body structure; the tentacles are simple and unbranched and normally number more than eight.

(2025). 9780520239395, University of California Press. .
These organisms are formed of individual soft polyps which in some species live in colonies and can secrete a skeleton. As with all , these organisms have a complex life cycle including a motile phase and a later characteristic sessile phase. Hexacorallia also include the significant extinct order of .

The taxon Hexacorallia is considered to be , that is all contained species are descended from a common ancestor.


Sea anemones
Certain or all -less hexacorals are called sea anemones in the literature. More specifically, sea anemones can mean one of the following:


Taxonomy

Phylogenetic classification
The current system is shown below:

class (or subclass) Hexacorallia aka::DeBiasse, M. B., Buckenmeyer, A., Macrander, J., Babonis, L. S., Bentlage, B., Cartwright, P., … Ryan, J. (2024). A Cnidarian Phylogenomic Tree Fitted With Hundreds of 18S Leaves. Bulletin of the Society of Systematic Biologists, 3(2). Https://academic.oup.com/sysbio/article/70/4/635/6122449?login=false Scrutton, Colin. (1997). The Palaeozoic corals, I: Origins and relationships. Proceedings of The Yorkshire Geological Society - PROC YORKS GEOL SOC. 51. 177-208. [4] Https://doi.org/10.17161/dt.v0i0.5541< /ref>

(2020). 9780081029091, Academic Press. .
Fautin, Daphne G., Sandra L. Romano, and William A. Oliver, Jr. 2000. Zoantharia. Sea Anemones and Corals. Version 04 October 2000. http://tolweb.org/Zoantharia/17643/2000.10.04 in The Tree of Life Web Project, accessed 2025-07-22
(2023). 9783031168871, Springer Nature. Https://www.scribd.com/document/676154745/Corals-and-Reefs-From-the-Beginning-to-an-Uncertain-Future%2523content%253Dquery:Coralomorpha,pageNum:97,indexOnPage:0&pg=PA87" title="Bar code 9783031168871 reference." target="_external">.
(1983). 9780521248563, CUP Archive. . (Note esp. the statement that "we consider that the Paleozoic "scleractiniamorphs" ... should be reclassified as genuine scleractinians")SEPKOSKI, J. J. A compendium of fossil marine animal genera. Bulletins of American palaentology. September 16, 2002 [5]
(2025). 9788478408054, IGME. .
(2009). 9781139478922, Cambridge University Press. .
  • order - tube anemones aka: (This order is sometimes placed outside Hexacorallia as a separate (sub)class.)
  • unnamed clade (i. e. Zoanthiniaria and related taxa):
    • order aka: – zoanthids aka:
    • ?† - coralomorphs (This is a semi-formal grouping of problematic genera, which in a broader sense also includes the below mentioned Tabulaconida and Cothoniida, as well as the †.)
    • order †
    • order †
    • order † (incl. †) – tabular corals aka: (In older systems, the Tabulata also included the , but these are now usually placed in . Moreover, the Tabulata were formerly sometimes placed outside Hexacorallia as a separate (sub)class.)
    • order † aka: (incl. †) – rugose corals aka: (The Rugosa were formerly sometimes placed outside Hexacorallia as a separate (sub)class.)
  • :
    • order (incl. †) – sea anemones aka:
    • order – corallimorpharians aka: (In palaentology, corallimorpharians are considered closely related to or even included in Actiniaria. But in texts dealing only with recent organisms, coralimorpharians are considered related to or even included in Scleractinia and the Corallimorpharia-Scleractinia clade is sometimes called . Corallimorpharia is not to be confused with Coralomorpha.)
    • order † (This is sometimes included in Scleractinia.)
    • order † (This is sometimes included in Scleractinia.)
    • order aka: (incl. †) – stony corals aka:
  • incertae sedis:
    • order – antipatharians aka: (This order probably belongs next to or inside Hexactiniaria.)
    • family
The order † (heterocorals), until recently considered related to or included in Rugosa, is now included in /ref>

Ceriantharia and Antipatharia were formerly jointly classified in a taxon called , which was considered a separate (sub)class outside Hexacorallia.


Skeleton-based classification
Based on the type of their skeleton, the hexacorals are divided into sea anemones, antipatharians and (Hexacorallian) corals. More specifically:
  • The , , , and have no , i. e. they are "soft", and are therefore called sea anemones. Note that the definition of the term sea anemone can vary (see the above chapter Sea anemones).
  • The have a horn-like - and therefore flexible - skeleton.
  • All the remaining (recent and fossil) taxa of the above system have a calcium carbonate - and therefore rigid ("hard") - skeleton, which, in this class, is always compact. They therefore constitute the main group of what is usually called (i. e. and with a compact rigid skeleton). Note that the definition of the term coral can vary, for example octocorals without a compact skeleton and Antipatharia are sometimes also included in the term corals.
In the 19th century, this skeleton-based division into three parts was also the standard formal way of subdividing the taxon hexacorals, i. e. the Hexacorallia aka consisted of the taxa (orders):
(2022). 9783368131807, BoD – Books on Demand. .
  • Actiniaria sensu lato aka:, i.e the sea anemones in the broadest sense;
  • Antipatharia aka:, i. e. the same as today's antipatharians;
  • Madreporaria sensu lato aka:, i. e. all the rest, meaning all Hexacorallian corals.

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