The Galleriinae are a subfamily of Pyraloidea (family Pyralidae) and occur essentially worldwide, in some cases aided by involuntary introduction by humans. This subfamily includes the , whose (waxworms) are bred on a commercial scale as food for and as fishing bait; in the wild, these and other species of Galleriinae may also be harmful to humans as pests.
At the species level, they are the least diverse snout moth subfamily according to current knowledge, with 306 described species all together. However, as regards major lineages, the Galleriinae are quite diverse, with five tribes being recognized – more than in the Phycitinae, the most species-rich snout moth subfamily. One of these tribes, the Joelminetiini, has been described only in 2007, and presently contains a single and highly aberrant genus.
Description and ecology
The
caterpillar of Galleriinae usually have a sclerotised (hardened) ring around the base of seta SD1 on the first abdominal segment. Their
are comparatively easy to distinguish from other snout moths' by a readily apparent midline ridge running along the thorax and abdomen. In the
imago, the
gnathos of the male
is reduced to the point of disappearing altogether or (more rarely) with only the barest vestige remaining; this is quite characteristic except for a few
Chrysauginae which have convergently lost the gnathos. Males produce very high chirping sounds with their tegulae, in some cases even regular "mating songs", though without specialized
bioacoustics equipment this cannot be used for identification. Unusual for Pyralidae, adult Galleriinae may lack ocelli and even the proboscis (which is usually well developed in the family); as typical for the family, however, they usually have large labial palps which form a "snout".
[Jia et al. (2001), Solis (2007), Zhou et al. (2008)]
, the subfamily is noted for a number of species that coevolved with Hymenoptera, namely Apoidea ( and relatives). The larvae may be or , and the adults of such species at least to some degree are (though usually pursued by the nest inhabitants). Especially notable among these Galleriinae are the ( Achroia and Galleria larvae) which are both significant as beekeeping pests and as commercial items, as well as Aphomia species. Others, especially the rice moth ( Corcyra cephalonica) and Paralipsa, are noted pests of stored food products.[Solis (2007)]
Systematics
The diversity of Galleriinae known in the mid-20th century was fully catalogued by P.E.S. Whalley of the UK Natural History Museum, but no dedicated
phylogenetic analysis has been conducted. Notwithstanding, such studies exist for the
Pyraloidea as a whole, and these indicate that the Galleriinae are a rather primitive lineage of Pyralidae, comparable to the
Chrysauginae. The latter may be the closest living relatives of the Galleriinae, or an independent but equally ancient snout moth lineage that simply looks similar due to sharing many
traits.
The Galleriinae are currently divided into five tribes, though this may change eventually. For one thing, some genera are presently insufficiently studied or too aberrant to be firmly assignable to any one tribe; for another, in the absence of detailed phylogenetic studies the best systematics treatment for members of this subfamily remains a best guess. Furthermore, not all Galleriinae are known to science; new species and genera continue to be discovered. A supposed additional tribe ("Macrothecini") is based on a misidentified specimen of Cacotherapia interalbicalis; on the other hand the genus Joelminetia, of which the first specimens reached the hands of researchers only in the 1990s, turned out to be so distinct as to warrant establishment of its own monotypic tribe.[Pitkin & Jenkins (2004), Solis (2007), and see references in Savela (2011)]
The tribes and genera – with some significant species also noted – in this subfamily are:[See references in Savela (2011)]
Cacotherapiini Munroe, 1995 (= Macrothecini)
Galleriini Zeller, 1848
Joelminetiini Speidel & Witt, 2007
Megarthridiini Whalley, 1964
Tirathabini Whalley, 1964
-
Acracona
-
Acyperas
-
Antiptilotis Meyrick, 1897
-
Aphomia
-
Bapara Walker, 1865
-
Callionyma Meyrick, 1882
-
Ceratothalama Meyrick, 1932
-
Rice moth – rice moth
-
Cristia Whalley, 1964
-
Doloessa Zeller, 1848
-
Eldana
-
Epimorius Zeller, 1877
-
Ertzica Walker, 1866
-
Ethopia Walker, 1865
-
Galleristhenia Hampson, 1917
-
Heteromicta Meyrick, 1886
-
Hypolophota Turner, 1904
-
Lamoria
-
Mampava Ragonot, 1888
-
Mecistophylla Turner, 1937
-
Metaraphia Hampson in Ragonot, 1901
-
Meyriccia Hampson, 1917
-
Microchlora Hampson in Ragonot, 1901
-
Neoepimorius Whalley, 1964
-
Neophrida Möschler, 1882
-
Paralipsa Butler, 1879
-
Paralipsa gularis – stored nut moth
-
Paraphomia Hampson in Ragonot, 1901
-
Parazanclodes Hampson in Ragonot, 1901
-
Paroxyptera Ragonot, 1901
-
Picrogama Meyrick, 1897
-
Pocopaschia Dyar, 1914
-
Pogrima Schaus, 1940
-
Prasinoxena Meyrick, 1894
-
Proropoca Hampson, 1916
-
Prosthenia Hampson in Ragonot, 1901
-
Schistotheca Ragonot, 1882
-
Statia Ragonot, 1901
-
Stenachroia Hampson, 1898
-
Stenopaschia Hampson, 1906
-
Thalamorrhyncha Meyrick, 1933
-
Tirathaba Walker, 1864
-
Xenophasma Dognin, 1905
incertae sedis
Footnotes
External links
