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Fynbos (; , ) is a small belt of natural shrubland or heathland vegetation located in the Western Cape and Eastern Cape provinces of South Africa. The area is predominantly coastal and mountainous, with a Mediterranean climate. The fynbos ecoregion is within the Mediterranean forests, woodlands, and scrub biome. In fields related to biogeography, fynbos is known for its exceptional degree of biodiversity and endemism, consisting of about 80% (8,500 fynbos) species of the Cape floral kingdom, where nearly 6,000 of them are endemic. The area continues to face severe human-caused threats, but due to the many economic uses of the fynbos, conservation efforts are being made to help restore it.
Of the world's six , this is the smallest and richest per unit of area. The Holarctic kingdom, in contrast, incorporates the whole of the Northern Hemisphere north of the tropics. The diversity of fynbos plants is extremely high, with over 9,000 species of plants occurring in the area, around 6,200 of which are endemic, i.e. growing nowhere else in the world. South Africa's Western Cape has the vast majority of species with one estimate finding 8,550 species in 89,000 km2, which is higher than that estimated for the Malayan forests, 7,900 species in 132,000 km2. It has been claimed that it exceeds even the richest tropical rainforest in South America, including the Amazon. Of the Ericas, over 600 occur in the fynbos kingdom, while only two or three dozen have been described in the rest of the world. This is in an area of 46,000 km2 – by comparison, the Netherlands, with an area of 33,000 km2, has 1,400 species, none of them endemic. Table Mountain in Cape Town supports 2,200 species, more than the entire United Kingdom. Thus, although the fynbos covers only 6% of the area of southern Africa, it has half the species on the subcontinent – and in fact has almost one in five of all African plant species so far described.
Five main river systems traverse the Cape floral kingdom: the Oliphants River of the Western Cape; the Berg River which drains the West Coast Forelands plain stretching from the Cape Flats to the Olifants; the Breede River, which is the largest river on the Cape; the Olifants River (Southern Cape); Gourits River and the Groot Rivers which drain the Little Karoo basin and the South Coast Forelands; and the Baviaanskloof and to the east.
Depending on the locality and the aspects under discussion, several other families have equal claim to being characteristic, including Asteraceae, Rutaceae, and Iridaceae. More than 1400 bulb species occur among the fynbos, of which 96 are Gladiolus and 54 Lachenalia. Areas that are dominated by "renosterbos", Dicerothamnus rhinocerotis, (Asteraceae) are known as Renosterveld (Afrikaans for "rhinoceros field").
The Lowland Fynbos and Renosterveld experiences regular winter rainfall, especially to the west of Cape Agulhas. The ecoregion has been subdivided into nine areas: the West Coast Forelands from the Cape Flats to the Olifants River (Western Cape); the Warm Bokkeveld basin around the town of Ceres; the Elgin Valley around the town of Elgin; the sandy Agulhas Plain on the coast; the Breede River valley around the town of Worcester; the South Coast Forelands from Caledon west to Mossel Bay; the south-eastern end of the Little Karoo; Langkloof valley; and the Southeastern Coast Forelands west from Tsitsikamma to Gqeberha.
The flora of the lowlands contains a high number of endemic species, and tends to favour larger plants than those growing on the hillier areas. They include the larger Restionaceae such as species of Elegia, Thamnochortus, and Willdenowia and proteas such as king protea ( Protea cynaroides) and blushing bride ( Serruria florida). Particular types of lowland fynbos include the shrubs and herbs of the coastal sand dunes, the mixture of ericoids and restoids with thickets of shrubs such as Maytenus, and other Celastraceae, and other Sapotaceae, and Rhus and other Anacardiaceae on the coastal sands; the classic fynbos of the sandplains of the West Coast Forelands, and the Agulhas Plain; the grassy fynbos of the hillier and wetter areas of the South and South-Eastern Coast Forelands; areas where fynbos and renosterveld are mixed; coastal renosterveld on the West and South Coast Forelands; and the inland renosterveld of the drier inland Little Karoo and Warm Bokkeveld.
The area is also home to a large number of endemic creatures that have adapted to life in this area, such as the which pollinate Ixia viridiflora. Endemic species of fish in the five river systems occur in the area, too. Endemic reptiles and amphibians include a number of and the chameleon-like arum frog ( Hyperolius horstockii).
The Montane Fynbos and Renosterveld is the area above , a total of of the Cape Fold Mountains. The same level of floral variety, including all three characteristic fynbos families, is found there, but ericas predominate. Because the higher and wetter areas are more protected and contain important water sources, the original flora is more intact than in the lowlands; but agriculture and global warming are still threats. The region includes the mountains in the west from the Cape Peninsula to the Kouebokkeveld Mountains, the south coast hinterland from Elgin to Gqeberha, the mountains north of the Little Karoo from Laingsburg to Willowmore, and the inselberg hills within the Little Karoo. About half of these areas are originally fynbos, and about half are renosterveld.
Many different microclimates occur, so the flora changes from west to east, and also varies with altitude up the hillsides away from the coast and according to compass direction. Lower elevations are covered with protea fynbos, with ericas taking over further up. Plant species include pincushions ( Leucospermum). The wildlife includes a number of endemic bees, beetles, horseflies, and ants, and birds such as and the orange-breasted sunbird. Many of these birds and insects are important and specific pollinators for the fynbos, such as the mountain pride butterfly ( Aeropetes tulbaghia) which only visits red flowers such as Disa uniflora and pollinates 15 different species. Larger animals include antelopes, particularly Cape grysbok ( Raphicerus melanotis), common duiker ( Sylvicapra grimmia), and klipspringer ( Oreotragus oreotragus). The extinct Bluebuck and quagga were also fynbos natives.
In many areas with Mediterranean climates, fynbos species have become popular garden plants, in particular and pelargonium, and in cooler regions are used as window plants.
A very large number of fynbos plant species are used in traditional medicine, and while only a tiny proportion have as yet been subjected to formal testing, many have already been identified as having medicinal properties.
The Working for Water (WfW) program was started in 1995 by the Department of Water Affairs and Forestry to control these invasive species which were shown to sequester 9.95% of usable surface water runoff. Since then, over 100,000 hectares of land have been cleared of invasive species while providing jobs to around 20,000 people per year, most of which are women and unskilled workers. Systematic monitoring of WfW's progress is lacking but there is anecdotal evidence that endemic silver peas have returned to Table Mountain after being thought extinct.
Vegetation types
Fynbos vegetation types, code FF:
Fauna
The fynbos is home to many unique and endemic animals, with seven species of endemic bird and an unknown number of endemic reptiles, amphibians, and arthropods. The seven avian endemics include the Cape rockjumper, Cape sugarbird, Victorin's warbler, Orange-breasted sunbird, Protea canary, Cape siskin, and Fynbos buttonquail.
Ecoregions
The fynbos area has been divided into two very similar : the lowland fynbos (below 300 m above sea level) on the sandy soil of the west coast, and the montane fynbos of the Cape Fold Belt.
Economic uses
Rooibos ( Aspalathus linearis) and honeybush ( Cyclopia intermedia) are of economic importance, grown and harvested in large quantities in the Cederberg area, and providing important exports. Restios continue to be used for thatching, as they have for hundreds or even thousands of years. Proteas and other floral species are grown in many areas and their flowers harvested for export.
Threats and conservation
The fynbos is the region of South Africa most affected by Invasive species which collectively cover around 10% of the entire country. The most common invasive plants are wattles and , native to Australia, and Pinus radiata native to Europe and the Californian coast of the United States. Pines had been introduced to South Africa by the 19th century and the wattles were imported into the mid-1870s to stabilize sand dunes. In 1997 it was estimated that invasion caused the fynbos region to decline in value by US$750 million per year.
See also
External links
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