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The Douglas fir ( Pseudotsuga menziesii) is an in the pine family, . It is the tallest tree in the Pinaceae family. It is to western and is also known as Douglas-fir, Douglas spruce, Oregon pine, and Columbian pine.

(2008). 9780470113691, John Wiley & Sons.
There are three varieties: coast Douglas-fir ( P. menziesii var. menziesii), Rocky Mountain Douglas-fir ( P. menziesii var. glauca) and Mexican Douglas-fir ( P. menziesii var. lindleyana).

Despite its , it is not a true ( Abies), (genus Picea), or (genus Pinus). It is also not a (genus Tsuga); the genus name Pseudotsuga means "false hemlock".


Description
Douglas-firs are medium-sized to extremely large trees, tall (although only coast Douglas-firs reach heights near 100 m) and commonly reach in diameter, although trees with diameters of almost exist. The largest coast Douglas-firs regularly live over 500 years, with the oldest specimens living for over 1,300 years. Rocky Mountain Douglas-firs, found further to the east,
(2025). 9781680513295, Mountaineers Books. .
are less long-lived, usually not exceeding 400 years in age.

There are records of former coast Douglas-firs exceeding in height, which if alive today would make it the tallest tree species on Earth. Particular historical specimens with heights exceeding include the Lynn Valley Tree and the .

The are flat, soft, linear needles long,

(2025). 9781604692631, . .
generally resembling those of the firs, occurring singly rather than in fascicles; they completely encircle the branches, which can be useful in recognizing the species. As the trees grow taller in denser forest, they lose their lower branches, such that the foliage may start as high as off the ground. Douglas-firs in environments with more light may have branches much closer to the ground.

The bark on young trees is thin, smooth, gray, and contains numerous blisters. On mature trees, usually exceeding 80 years, it is very thick and corky, growing up to thick with distinctive, deep vertical fissures caused by the gradual expansion of the growing tree. Some of the mature bark is brown, while other parts are lighter colored with a cork-like texture; these develop in multiple layers. This thick bark makes the Douglas-fir one of the most fire-resistant tree native to the Pacific Northwest.

The male are yellowish red, long. The female cones are green when young, maturing to reddish-brown or gray, long; they are pendulous, with persistent scales, unlike those of true firs. They have distinctive long, trifid (three-pointed) which protrude prominently above each scale and are said to resemble the back half of a mouse, with two feet and a tail. The seeds are long, with a longer wing.

The massive of Douglas-fir was sequenced in 2017 by the large PineRefSeq consortium, revealing a specialized photosynthetic apparatus in the light-harvesting complex of genes.


Taxonomy
The common name honors David Douglas, a Scottish botanist and collector who first reported the extraordinary nature and potential of the species. The common name is misleading since it is not a true fir, i.e., not a member of the genus . For this reason, the name is often written as Douglas-fir (a name also used for the genus as a whole). Douglas' connection to the Douglas fir is told in the 1962 Death Valley Days episode 'Grass Man'.

The specific epithet menziesii is after Archibald Menzies, a Scottish physician and rival to David Douglas. Menzies first documented the tree on in 1791. Colloquially, the species is also known simply as Doug fir or Douglas pine (although the latter common name may also refer to Pinus douglasiana). Other names for this tree have included Oregon pine, British Columbian pine, Puget Sound pine, Douglas spruce, false hemlock, red fir, or red pine (although again red pine may refer to a different tree species, , and red fir may refer to ).

One name for the tree, used in the language, is italic=yes. In the language, the tree is called italic=yes.


Distribution
Pseudotsuga menziesii var. menziesii, the coast Douglas-fir, grows in the coastal regions from west-central southward to Central California. In and Washington, its range is continuous from the eastern edge of the west to the Pacific Coast Ranges and Pacific Ocean. In California, it is found in the Klamath and California Coast Ranges as far south as the Santa Lucia Range, with a small stand as far south as the in Santa Barbara County. One of the last remaining old growth stands of conifers is in the Mattole Watershed, and is under threat of logging. In the , it ranges as far south as the region. It occurs from sea level along the coast to elevations of or higher, and inland in some cases up to .

Another variety exists further inland, Pseudotsuga menziesii var. glauca, the Rocky Mountain Douglas-fir or interior Douglas-fir. Interior Douglas-fir intergrades with coast Douglas-fir in the Cascades of northern Washington and southern British Columbia, and from there ranges northward to central British Columbia and southeastward to the Mexican border, becoming increasingly disjunct as latitude decreases and altitude increases. Mexican Douglas-fir ( P. lindleyana), which ranges as far south as , is often considered a variety of P. menziesii.

Fossils (wood, pollen) of Pseudotsuga are recorded from the and of Europe (, , Austria).

It is also naturalised throughout Europe, Argentina and Chile (called Pino Oregón). In New Zealand it is considered to be an invasive species, called a , and is subject to control measures. But is also one of the most common lumber trees used in forestry alongside Radiata pine with large plantations throughout the country. The species was introduced in the 20th century for its wood.


Ecology

Preferred sites
Douglas-fir prefers acidic or neutral soils, such as Olympic soil. However, it exhibits considerable morphological plasticity, and on drier sites P. menziesii var. menziesii will generate deeper taproots. Pseudotsuga menziesii var. glauca exhibits even greater plasticity, occurring in stands of interior temperate rainforest in British Columbia, as well as at the edge of semi-arid sagebrush steppe throughout much of its range, where it generates even deeper taproots still.

The coast Douglas-fir variety is the dominant tree west of the Cascade Mountains in the Pacific Northwest. It occurs in nearly all forest types and competes well on most parent materials, aspects, and slopes. Adapted to a more moist, mild climate than the interior subspecies, it grows larger and faster than Rocky Mountain Douglas-fir. Associated trees include , , , western white pine, , , , , , Lawson's cypress, , and several others. Pure stands are also common, particularly north of the in Oregon. It is most dominant in areas with a more frequent fire regime that suppresses less fire-resistant conifers.


Use by animals
Douglas-fir seeds are an extremely important food source for small mammals such as moles, , and , which consume an estimated 65% of each annual seed crop. The harvests and hoards great quantities of Douglas-fir cones, and also consumes mature pollen cones, the inner bark, terminal shoots, and developing young needles.

Mature or "old-growth" Douglas-fir forest is the primary habitat of the red tree vole ( Arborimus longicaudus) and the ( Strix occidentalis). Home range requirements for breeding pairs of spotted owls are at least of old growth. Red tree voles may also be found in immature forests if Douglas-fir is a significant component. The red vole nests almost exclusively in the foliage of the trees, typically above the ground, and its diet consists chiefly of Douglas-fir needles.

Douglas-fir needles are generally poor browse for , although in the winter when other food sources are lacking it can become important, and black-tailed deer browse new seedlings and saplings in spring and summer. The spring diet of the features Douglas-fir needles prominently.

The leaves are also used by the ; this 0.5 mm-long sap-sucking is conspicuous on the undersides of the leaves by the small white "fluff spots" of protective wax that it produces. It is often present in large numbers, and can cause the foliage to turn yellowish from the damage it causes. Exceptionally, trees may be partially defoliated by it, but the damage is rarely this severe. Among , apart from some that feed on Pseudotsuga in general, the Chionodes abella and C. periculella as well as the cone scale-eating have been recorded specifically on P. menziesii.

The inner bark is the primary winter food for the North American porcupine.

is a flavanone, a type of flavonoid, produced by P. menziesii in reaction to infection by .


Value to other plants
A which uses P. menziesii is the Douglas-fir dwarf mistletoe ( Arceuthobium douglasii). such as and mosses are common sights on Douglas-firs. As it is only moderately , undisturbed Douglas-fir stands in humid areas will eventually give way to later successional, more shade-tolerant associates such as the western redcedar and western hemlock—though this process may take a thousand years or more. It is more shade tolerant than some associated fire-dependent species, such as and ponderosa pine, and often replaces these species further inland.


Diseases and insects
Fungal diseases such as laminated root rot and shoestring root-rot can cause significant damage, and in plantation settings dominated by Douglas-fir monocultures may cause extreme damage to vast swathes of trees. Interplanting with resistant or nonhost species such as western redcedar and can reduce this risk. Other threats to Douglas-fir include and the Douglas-fir beetle.


Uses
Many different Native American groups used the bark, resin, and needles to make herbal treatments for various diseases. built waʻa kaulua () from coast Douglas-fir logs that had ashore. The wood has historically been favored as , especially from the coastal variety. In addition early settlers used Douglas-fir for all forms of building construction, including floors, beams, and fine carving.

The species is extensively used in management as a tree for softwood . Douglas-fir is one of the world's best timber-producing species and yields more timber than any other species in North America, making the forestlands of western Oregon, Washington, and British Columbia the most productive on the continent. In 2011, Douglas-fir represented 34.2% of US lumber exports, to a total of 1.053 billion (2,484,803,3 ). Douglas-fir timber is used for timber frame construction and timber trusses using traditional joinery, , and due to its strength, hardness and durability. As of 2024, the only wooden ships still currently in use by the U.S. Navy in conventional naval operations are Avenger-class minesweepers, made of Douglas-fir.

Douglas-fir sees wide use in heavy timber structures, as its wood is strong, available in a number of specifications including kiln dried and grade stamped, and can be supplied in very long lengths to . West coast mills are sophisticated in their processing of timbers, making lead times predictable and availability reliable. adhere well to Douglas-fir. perform well on Douglas-fir timbers with the mild caution that the natural color of this species varies and care must be taken to ensure uniformity of color. Pitch pockets that may ooze resin can be present in timbers that have not been kiln dried. Because of the timber sizes available, stamped timber grading, and relatively short lead times, Douglas-fir sees wide use in both public and residential projects.

The species has ornamental value in large parks and gardens. It has been commonly used as a since the 1920s, and the trees are typically grown on plantations.

The buds have been used to flavor eau de vie, a clear, colorless fruit brandy. Douglas-fir pine leaves can be used to make pine needle tea. They possess a tangy citrus flavor and may serve in some recipes as a wild substitute for .


Notable examples
The immense size and long lifespan of the Douglas fir has produced some acclaimed individual trees. Historical examples include the Lynn Valley Tree near Vancouver, British Columbia and the in in Washington State.

Living examples in Canada include the Red Creek Fir in the San Juan Valley of , the largest known Douglas-fir in Canada and one of the largest in the world; the Bonin Giant in the watershed in British Columbia's and Big Lonely Doug north of on ..

American examples include the Lake Quinault Douglas fir in the Olympic National Forest, Washington State, the largest known Douglas fir in the United States; Coast Douglasfir (Pseudotsuga menziesii) champion, American Forests, listed 2018-09-05; accessed 2019-01-05 the in North Fork Coquille River in , the second greatest of a living Douglas fir in the United States.; the , in the Olympic National Park in Washington State with the largest known in diameter; The Gymnosperm Database: Pseudotsuga menziesii subsp. menziesii, last modified 2018-12-23; accessed 2019-01-05 and the fir in Coos County, Oregon.

Outside of North America, the Hermitage Douglas-fir in , Scotland was one of the tallest trees in the until it fell in 2017 and the Waldtraut of the mill forest, in Freiburg im Breisgau is the tallest tree in Germany.


See also
  • List of Douglas-fir species
  • List of Douglas-fir diseases
  • List of superlative trees


Further reading
  • Brock, Emily K. (2015). Money Trees: The Douglas-Fi.r and American Forestry, 1900–1944 Corvallis, Oregon: Oregon State University Press.


External links

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