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   » » Wiki: Clupeidae
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Clupeidae is a family of ray-finned fishes, comprising, for instance, the and . Many members of the family have a body protected with shiny cycloid (very smooth and uniform) scales, a single , and a fusiform body for quick, evasive swimming and pursuit of prey composed of . Due to their small size and position in the lower of many marine food webs, the levels of they are very low, reducing the risk of mercury poisoning when consumed.

The earliest known fossil members of this group are the stem-clupeids and from the late Campanian/early Maastrichtian of Italy.


Description and biology
Clupeids are mostly marine , although a few species are found in fresh water. No species has scales on the head, and some are entirely scaleless. The is short or absent, and the teeth are unusually small where they are present at all. Clupeids typically feed on , and range from in length.

Clupeids spawn huge numbers of eggs (up to 200,000 in some species) near the surface of the water. After hatching, the larvae live among the plankton until they develop a and transform into adults. These eggs and fry are not protected or tended to by parents. The adults typically live in large shoals, seeking protection from predators such as , and other predatory fish, , marine mammals, and jellyfish. They also form .

(1998). 9780125476652, Academic Press.

Commercially important species of the Clupeidae include the ( Clupea harengus), and the ( C. pallasii).


Feeding physiology
The Clupeidae family primarily feed on small planktonic organisms. The teeth of members of this family are either reduced or absent, reduced teeth are miniature teeth that would be barely visible and line the interior of the fish's mouth. The structure of these teeth indicate that these organisms do not need to cut or tear their prey items as they would need fully formed teeth to complete this process. They do, however, possess long that are designed for sifting plankton and other small particles out of the water as it passes through their gills. Gill rakers are protrusions along the , opposing the gill filaments, that help aquatic organisms to trap food particles.

The diet of many clupeids primarily consists of and plant matter during their larval stages. As the fish mature this diet begins to shift towards larger and more substantive organisms, including more and . This change in diet is possible due to their increase in body and gill raker size, which allows them to capture and process larger organisms to support themselves. Small organisms like these do not need to be ground or torn apart for consumption so pronounced teeth would not serve a purpose in the feeding habits of Clupeidae, instead the use of allows for much more efficient nutrient collection.

The body shape of Clupeidae is also advantageous to their trophic ecology. The tapering body form is a highly hydrodynamic form that allows for quick increases in speed and a high maximum speed. Moving at high speeds allows the members of this family to regulate their feeding habits and avoid predators. Clupeidae can moderate the speed at which they swim to increase their uptake of nutrients. As with all filter feeders, Clupeidae cannot take in food if nutrient rich water does not pass over their gills. To moderate this, members of this family have been found to increase their swimming speed when they sense that there is a high concentration of food items in order to take advantage of this feeding period. Keeping a high swimming speed during periods of low food availability would not be efficient to maintain over long periods of time as the organisms would not net as much energy as they may need to in order to sustain themselves and increase their fitness. Increasing their swimming speed during feeding periods would allow them to take in more plankton while not suffering consequences from maintaining that speed.


Taxonomy
The following genera are classified within the family:

The family arguably also contains the "", a taxon first thought to be a distinct family, but then discovered to be deeply nested in the Clupeidae.

Until recently, the concept of Clupeidae was broader, but it has been subdivided into several distinct families (e.g. )


Fossil genera
The following fossil genera have been variously suggested to be sensu stricto members of Clupeidae. Many were formerly placed in the subfamily Clupeinae:


Disputed fossil genera
Known fossil genera classified under the concept of Clupeidae include:

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