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Cannabaceae is a small family of , known as the hemp family. As now circumscribed, the family includes about 170 species grouped in about 11 genera, including (hemp), () and (hackberries). Celtis is by far the largest genus, containing about 100 species.Stevens, P.F. (2001 onwards) "Cannabaceae", Angiosperm Phylogeny Website, retrieved 2014-02-25

Cannabaceae is a member of the . Members of the family are erect or climbing plants with petalless flowers and dry, one-seeded fruits. Hemp ( Cannabis) and hop ( Humulus) are the most economically important species.

Other than a shared evolutionary origin, members of the family have few common characteristics; some are trees (e.g. Celtis), others are (e.g. Cannabis).


Description
Members of this family can be trees (e.g. ), erect herbs (e.g. ), or twining herbs (e.g. ).

Leaves are often more or less palmately lobed or palmately compound and always bear . are always present and some members of this family possess .

Cannabaceae are often (distinct male and female plants). The flowers are (radially symmetrical) and not showy, as these plants are . As an adaptation to this kind of pollination, the calyx and corolla are radically reduced to only vestigial remnants found as an adherent perianth coating the seed. A reduced and monophyllous cuplike perigonal bract, properly known as the bracteole, immediately surrounds and protects the seed and is often misnamed as a "calyx". Flowers are grouped to form cymes. In the dioecious plants the male inflorescences are long and look like , while the female ones are shorter and bear fewer flowers. The is made of two connate , the usually superior ovary is unilocular; there is no fixed number of .

The fruit can be an or a .


Taxonomy

Classification
Classification systems developed prior to the 1990s, such as those of Cronquist (1981) and Dahlgren (1989), typically recognized the order , which included the families Cannabaceae, Cecropiaceae, Celtidaceae, Moraceae, Ulmaceae and Urticaceae, as then circumscribed. Molecular data from 1990s onwards showed that these families were actually embedded within the order Rosales, so that from the first classification by the Angiosperm Phylogeny Group in 1998, they were placed in an expanded Rosales, forming a group which has been called 'urticalean rosids'.'' with nearly mature flowers ()]] Cannabaceae comprises the following genera:

  • Trema Lour. (12–42 spp.)


Phylogeny
Cannabaceae likely originated in East Asia during the Late . The oldest known pollen typical of members of Cannabaceae is from the Late Cretaceous ( ~94–90 million years ago) of , Borneo. Fossils show Cannabaceae were widely distributed in the Northern Hemisphere during the early Cenozoic, though their distribution shifted towards tropical regions in the later Cenozoic due to changing climates.

Modern molecular phylogenetics suggest the following relationships:


Uses
has revealed that these plants may have been used for ritual/medicinal purposes in Xinjiang, China as early as 494 B.C.

, the common hop, has been the predominant bittering agent of for hundreds of years. The flowers' resins are responsible for beer's bitterness and their ability to extend shelf life due to some qualities. The young shoots can be used as a vegetable.

Some plants in the genus Cannabis are cultivated as for the production of fiber, as a source of cheap , for their nutritious seeds, or their edible leaves. Others are cultivated for medical or recreational use as dried flowers, , or . Induced in pistilate flowers, and are used to produce either higher or lower yields of tetrahydrocannabinol (THC), other , as well as with desired flavors or aromas, such as blueberry, strawberry, or even citrus.

Many trees in the genus Celtis are grown for landscaping and ornamental purposes, and the bark of is used to produce high-end Chinese paper.


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