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The Cabombaceae are a family of aquatic, herbaceous . A common name for its species is water shield. The family is recognised as distinct in the Angiosperm Phylogeny Group IV system (2016). The family consists of two genera of , and , totalling six species.


Description

Vegetative characteristics
Cabombaceae are perennial, rhizomatous, aquatic herbsCabombaceae in Flora of North America @ efloras.org. (n.d.). Retrieved February 17, 2025, from http://www.efloras.org/florataxon.aspx?flora_id=1&taxon_id=10140 with slender, branched rhizomes and adventitious roots. The leaves are whorled, alternate, or opposite. Both floating and submerged leaves are produced. Https://vicflora.rbg.vic.gov.au/flora/taxon/0fbf1892-c184-4ba6-8217-f627ab8fb0ec< /ref>

Generative characteristics
The solitary, pedunculate, bisexual, chasmogamous, actinomorphic, inodorous Https://floradobrasil.jbrj.gov.br/FB69< /ref> flowers float on the water surface or extend beyond it.Cabombaceae in Flora of China @ efloras.org. (n.d.). Retrieved February 17, 2025, from http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=10140 The gynoecium consists of (1–)2–18 free carpels. Https://profiles.ala.org.au/opus/foa/profile/Cabombaceae Date The indehiscent, Https://www.biodiversitylibrary.org/page/25672588< /ref> follicle-like or achene-like fruit bears 1–3 seeds.


Distribution
The Cabombaceae are all aquatic, living in still or slow-moving waters of temperate and tropical North and South America, Europe, Asia, Africa, and Australia. Although found on all continents but Antarctica, the plants tend to grow in relatively restricted ranges.


Fossil record
The family has an extensive fossil record from the with plants that exhibit affinities to either the Cabombaceae or Nymphaeaceae occurring in the Early Cretaceous.

One such likely Cretaceous member is the genus , found in rocks 115 million years old in what is now Brazil. David W.Taylor, G.J.Brenner et S.H.Basha has been described from the Lower Cretaceous of .Taylor, D. W., Brenner, G. J., & Basha, S. D. H. (2008). " Scutifolium jordanicum gen. et sp. nov.(Cabombaceae), an aquatic fossil plant from the Lower Cretaceous of Jordan, and the relationships of related leaf fossils to living genera. American Journal of Botany, 95(3), 340-352. Krassilov et Bachia has been described from the mid-Cretaceous of Morocco.


Taxonomy

Publication
The family Cabombaceae Rich. ex A.Rich. was first published in 1822 by Louis Claude Richard, but initial description did not satisfy the requirements for valid publications. The family then was validated by .


Taxonomic history
The of 1998 included this family in the water lily family , as did the APG II system, of 2003 (optionally). The APG III and APG IV systems of classification separated the family Cabombaceae from the family Nymphaeaceae. The family is part of the order Nymphaeales, which is one of the most basal lineages.


Further reading
  • Simpson, M.G. Plant Systematics. Elsevier Academic Press. 2006.


External links
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