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Caatinga () is a type of tropical vegetation, and an characterized by this vegetation in interior northeastern Brazil. The name "Caatinga" comes from the word , meaning "white forest" or "white vegetation" ( = forest, vegetation, = white). The Caatinga is a xeric shrubland and , which consists primarily of small, thorny trees that shed their leaves seasonally. , thick-stemmed plants, thorny brush, and arid-adapted grasses make up the ground layer. Most vegetation experiences a brief burst of activity during the three-month-long .

Caatinga falls entirely within earth's and is one of six major biomes of Brazil. It covers 912,529 km², nearly 10% of Brazil's territory. It is home to 26 million peopleSalcedo, I.H., Menezes, R.S.C. (2009): Agroecosystem functioning and management in semi-arid Northeastern Brazil, in: Tiessen, H., Stewart, J.W.B. (eds.): Applying Ecological Knowledge to Landuse Decisions. Inter-American Institute for Global Change Research - IICA-IAI-Scope, Paris, pp. 73–81. and over 2000 species of plants, fish, reptiles, amphibians, birds, and mammals.

The Caatinga is the only exclusively Brazilian , which means that a large part of its biological heritage cannot be found anywhere else on the planet.


Geography
The Caatinga covers the interior portion of northeastern Brazil bordering the Atlantic seaboard (save for a fringe of ), extending across nine states: Piauí, Ceará, Rio Grande do Norte, Paraíba, , , , , and parts of . Altogether, the Caatinga comprises 850,000 km², about 10% of the surface area of Brazil. By comparison, it is over nine times the surface area of , whence came Brazil's early European settlers.

Located between 3°S 45°W and 17°S 35°W, the Caatinga experiences irregular winds from all directions. Rainfall is thus intermittent but intense, totalling on average. Although the climate is typically hot and semi-arid, the Caatinga includes several enclaves of humid tropical forest, with trees tall.

To the northwest, the Caatinga is bounded by the Maranhão Babaçu forests; to the west and southwest, the Atlantic dry forests and savannas; to the east, the humid ; and to the north and northeast, the Atlantic Ocean.


Climate
During the dry winter periods there is no or undergrowth, as plants try to conserve water. Roots protrude through the surface of the stony soil, to absorb water before it is evaporated. Leaves fall off the trees to reduce . With all the foliage and undergrowth dead during the drought periods and all the trees having no leaves the Caatinga has a yellow-grey, -like look. During the peak periods of drought the Caatinga's soil can reach temperatures of up to 60 °C.

The drought usually ends in December or January, when the rainy season starts. Immediately after the first rains, the grey, desert-like landscape starts to transform and becomes completely green within a few days. Small plants start growing in the now moist soil and trees grow back their leaves. Rivers that are mostly dry during the past six or seven months start to fill up and streams begin to flow again.


Ecology
Caatinga harbors a unique biota, with thousands of endemic species. Caatinga contains over 1,000 vascular plant species in addition to 187 bees, 240 fish species, 167 reptiles and amphibians, 516 birds, and 148 mammal species, with endemism levels varying from 9 percent in birds to 57 percent in fishes.


Vegetation
The Caatinga does not correspond to a single type of vegetation, but rather a broad mosaic. Nonetheless, all vegetative structure is adapted to the xeric climate. and species dominate; non-succulents exhibit small, firm leaves and intense branching at the base, akin to shrubs. Palm stands usually contain or babaçu palms, but occasionally tucumã and macaúba.

The Caatinga has enough endemic species to constitute a floristic province.

Most authors divide the Caatinga into two different subtypes: dry ("sertão") and humid ("agreste"), but categorizations vary to as many as eight different vegetative regimes.


Fauna
The Caatinga is home to nearly 50 species of birds, including Lear's macaw (Anodorhynchus leari), Spix's macaw (Cyanopsitta spixii), moustached woodcreeper (Xiphocolaptes falcirostris), Caatinga parakeet, , Sao Francisco black tyrant and Caatinga cacholote.

Endemic mammal species include:


Indigenous peoples
The oldest human remains in the Caatinga are found in the Serra da Capivara National Park, in Piauí, where artifacts, rock paintings, and a skull named "Zuzu" were discovered, dating back approximately 8,000 years. This fossil, under study, may be even older than the Luzia fossil, which is currently the oldest in the South American continent.

The caatinga was inhabited by two major indigenous groups, the Macro-Jê and the Kariris, who have been in the Caatinga for at least two thousand years. After the 11th century, the Tupis arrived in the region, coming from the southeast and through the Atlantic coast. Contact with colonizers starting in the 16th century decimated numerous indigenous nations and tribes through diseases, enslavement, and invasion of territories for cattle ranching, sugar mills, and new settlements. Many of the Northeastern indigenous peoples chose assimilation, abandoning their customs, language, and religion to survive European advances, so many Northeasterners are mixed descendants of indigenous peoples and Europeans.

Currently, the Caatinga still has indigenous peoples, the largest of which are the Potyguaras, of Tupi origin and also native to the Atlantic Forest, totaling more than 20,000 indigenous peoples. In the interior, the largest groups are the Xukurus and Pankarus, from the Pernambuco Caatinga, totaling 12,000 and 7,000 indigenous peoples, possibly of Macro-Jê origin. The Fulni-Ô people are known for being the only indigenous ethnic group in the Northeast to have kept their ancestral language alive, as well as having saved unique cultural elements such as the Ouricuri Ritual; they are one of the least acculturated Northeastern peoples by European invaders. Other notable Caatinga peoples are the Kambiwás, Tremembés, Pitaguarys, Kariris, Kiriris, and Tabajaras.


Possible anthropogenic origins
Based on radiocarbon dating of , proponents of historical ecology such as and William Balee have suggested that large sections of the Caatinga region may be of anthropogenic origin. Over 1000 years ago, native peoples may have unintentionally created the environment of the modern-day Caatinga through constant , thereby stymying plant succession and preventing major rainforests from growing within the region.
(2013). 9780817317867, University of Alabama Press. .

Conversely, fossil evidence suggests that the Caatinga may historically have been part of a much larger dry belt.


Conservation
The Caatinga is poorly represented in the Brazilian Conservation Area network, with only 1% in Integral Protection Conservation Areas and 6% in Sustainable Use Conservation Areas. Protected areas include Chapada Diamantina National Park, Serra da Capivara National Park, and Serra das Confusões National Park.

Economic developed has fragmented the native biome. Estimates on the amount of Caatinga transformed affected by economic development range 25-50%, making Caatinga the most degraded ecosystem in Brazil, following the Atlantic Forest, which has lost over 80% of its original cover.


Economic exploitation
The local population lives in extreme poverty, and many rely on extraction of natural resources for a livelihood. There are few drinkable water sources, and harvesting is difficult because of the irregular rainfall.


Agriculture
Native plants are used in local agriculture, much of it . Pilocarpus jaborandi appears to exhibit medicinal properties. The fruits of umbú and are used as food directly, and other species are used for forage. Local palms produce commercial-grade and oils, which undergirds much of the economy of northeast Brazil.

is also a well-developed and traditional activity in the region. One of the most productive species, Melipona subnitida, known locally as jandaíra, produces up to 6 liters of honey a year, resulting in economic profit for the population.

Irrigation along the São Francisco River promises to turn the region into a breadbasket. The soil is very fertile, and existing irrigation infrastructure already supports the export of , and . At the same time, irrigation threatens to salinize the soil.


Grazing
(Guzerá and cattle) and farming are popular and very productive in the region. and timbering for fuelwood have decimated local vegetative populations; outside irrigated regions, the area has begun to à la Sahara and Sahel.


See also

On Caatinga
  • Caatinga moist-forest enclaves
  • Northeastern Brazil
  • Sertão
  • List of plants of Caatinga vegetation of Brazil


The five other major ecoregions of Brazil


Notes

Further reading
  • Vargas Llosa, Mario The War of the End of the World


Historical biogeographic surveys
  • Marcgrav (1638)
  • Spix & Martius (1817-1820)


External links

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