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   » » Wiki: Azendohsauridae
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Azendohsauridae is a family of that lived during the to period, around 242-216 million years ago. The family was originally named solely for the eponymous , marking out its distinctiveness from other allokotosaurs, but the family now includes four other : the basal genus , the large horned herbivore , and two carnivorous genera grouped into the -level subclade Malerisaurinae, and , and potentially also the dubious genus Otischalkia. Most fossils of azendohsaurids have a distribution, with multiple species known across and in as well as , although fossils of malerisaurine azendohsaurids have also been found in the southwestern of .

Azendohsaurids are notable for the various dinosaur-like traits found in some species, including the -like neck, jaws and teeth of Azendohsaurus, the -like horns of Shringasaurus, and -like teeth of Puercosuchus. These traits are all convergently evolved with later dinosaurs, and some similarities are so striking that it is difficult to distinguish isolated azendohsaurid teeth and jaw bones from those of dinosaurs. Indeed, Azendohsaurus itself was initially described as a herbivorous dinosaur until better remains of its skull and skeleton were found.


Description
Azendohsaurids were robust quadrupeds with sprawled limbs, characterised by their long necks and proportionately small heads, and reached body sizes of up to in length in the largest species. Although initially characterised as herbivores based solely on Azendohsaurus, azendohsaurids had a diverse diet and lifestyles including large herbivores, insectivores, and carnivores. Consequently, although broadly similar in body form, their skulls varied from short and boxy with leaf-shaped teeth, to long and narrow with recurved, blade-like teeth.

Azendohsaurids lack a prenarial process, the bony splint of the that otherwise divides the two in typical reptile skulls, giving them a single fused (confluent) opening for their nostrils. Another characteristic of azendohsaurids is that they often possess palatal teeth on the roof of their mouths similar in size and shape to those along the jaw margins, including serrations, compared to the typical tiny and conical palatal teeth of other reptiles. Such large and specialised palatal teeth are known definitively in Pamelaria, Azendohsaurus and Shringasaurus, while simpler but notably large palatal teeth with coarse serrations are found in Malerisaurus. The palatal teeth of Puercosuchus, however, are simply described as "peg-like".

All known azendohsaurids have long necks, with very similar in shape and construction to those of early sauropodomorph dinosaurs, and held them raised above their shoulders. The themselves are tall and very well developed, with long blades. Azendohsaurids had a sprawling gait like other early archosauromorphs, although their faces back as well as out to the sides, suggesting they may have been able to hold their forelimbs closer to their body. Their bodies are deep, ranging from relatively narrow-bodied in Pamelaria to barrel-shaped in Azendohsaurus, and their tails are proportionally shorter and stockier compared to other archosauromorphs.


Classification and evolution
Azendohsaurids are one of two or three families included in the clade Allokotosauria, a group of unusual Triassic non- archosauromorphs that also includes the families and possibly the gliding . They have consistently been recognised as the of trilophosaurids, initially united on shared yet differing herbivorous traits. However, as more azendohsaurids have been discovered and recognised, they demonstrate that the group was likely to be ancestrally carnivorous.

A phylogenetic analysis performed by palaeontologist Sterling J. Nesbitt and colleagues in 2021 included all recognised azendohsaurids and other allokotosaurian taxa. Their results found Pamelaria to be the earliest branching azendohsaurid, with the remaining azendohsaurids divided into two subclades, one containing the herbivores Shringasaurus and two species of Azendohsaurus, and the Malerisaurinae containing both species of Malerisaurus and similar larger material (including the material that would be named Puercosuchus). The below depicts the simplified consensus result of their analysis, where the uncertain relationships within Malerisaurinae are presented as a :


Biogeography
The oldest known azendohsaurids are Pamelaria and Shringasaurus, both from India and dated to the stage of the . Azendohsaurus itself has been dated from the end of the Middle Triassic during the (at least for A. madagaskarensis) into the stage of the earliest in Morocco and Madagascar. Malerisaurines, including Malerisaurus and Puercosuchus, are exclusively known from the during an interval from the late Carnian into the early of both India and North America, and are the latest-surviving azendohsaurids known. This is in spite of their relatively (i.e. ancestral) features compared to other azendohsaurids, representing a relictual lineage of early-diverging carnivorous azendohsaurids that survived longer than their more derived herbivorous kin. Malerisaurines disappear from the fossil record in North America at or the near the end of the (a local unit in the southwestern United States) roughly 216 million years ago, marking the extinction of the azendohsaurids globally.

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