The genus Aptenodytes contains two extant species of collectively known as "the great penguins".
Combined morphological and Molecular clock have shown the genus Aptenodytes to be basal to all other living penguins, that is, the genus split off from a branch which led to all other species. DNA evidence suggests this split occurred around 40 million years ago. This had been foreshadowed by an attempt to classify penguins by their behavior, which also predicted the genus' basal nature.
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