Antarctosaurus (; meaning "southern lizard") is a genus of sauropod dinosaur from the Late Cretaceous Period of what is now South America. The type species, Antarctosaurus wichmannianus, and a second species, Antarctosaurus giganteus, were described by prolific Germany paleontology Friedrich von Huene in 1929. Three additional species of Antarctosaurus have been named since then but later studies have considered them dubious or unlikely to pertain to the genus .
The type species, A. wichmannianus, is controversial because there is uncertainty as to whether all the described remains belong to the same individual or even genus. The second species , A. giganteus, is considered dubious, but the fragmentary remains represent one of the largest dinosaurs known.
Antarctosaurus wichmannianus is the type species of the genus, named in 1929 after the discoverer of its remains in 1912, geologist Ricardo Wichmann. Von Huene used the name A. wichmannianus to describe a large assemblage of bones, which are considered to come from the Anacleto Formation in Río Negro Province of Argentina, which is probably early Campanian in age. Two additional limb bones, found in the Chubut Province in 1924, were also referred to A. wichmannianus by von Huene in 1929. Later studies, however, have doubted their referral to the species.
Von Huene also named a fragmentary second species of Antarctosaurus in the same 1929 monograph, which he tentatively called cf. Antarctosaurus giganteus because of its enormous size. These fossils were recovered in Neuquén Province of Argentina, from the Plottier Formation, which dates to the Coniacian-Santonian stages of the Late Cretaceous Period. The Plottier, like the younger Anacleto, is a member of the Neuquén Group.
Very few remains are known of this species and it is regarded as a nomen dubium by some.Upchurch, P., Barrett, P.M, & Dodson, P. 2004. Sauropoda. In: Weishampel, D.B., Dodson, P., & Osmolska, H. (Eds.). The Dinosauria (2nd Edition). Berkeley: University of California Press. Pp. 259-322. Other researchers regard A. giganteus as a likely valid species but probably belonging to a new genus. In 1969, Leigh Van Valen considered A. wichmannianus and A. giganteus to be growth stages of the same species and favored the name A. giganteus. This idea is problematic because A. wichmannianus was named earlier in the same paper and it is known from more material, it should, therefore, get priority over A. giganteus. The two species are also not from the same geological formation which suggests they did not belong to the same time period.
In 1933, Von Huene and Charles Matley described another species, Antarctosaurus septentrionalis, meaning "northern". The remains were found in the Lameta Formation of Madhya Pradesh in India.
Antarctosaurus jaxarticus from Kazakhstan is known from a single femur.
In 1970, two fragmentary limb bones and a partial vertebra were found in the Adamantina Formation (originally described as Bauru Group; has also been reported as the São José do Rio Preto Formation) of the northern Paraná Basin in Brazil. The remains were described by their discoverers Fahad Moysés Arid and Luiz Dino Vizotto in 1971 as A. brasiliensis.Arid, F.M. & Vizotto, L.D. 1971. Antarctosaurus brasiliensis, um novo saurópode do Crétaceo superior do sul do Brasil. In: Congresso Brasileiro de Geologia 25: 297-305. In Other researchers have considered this species as either, a nomen dubium, or an indeterminate titanosaur.
The remains that have been described belong to Sauropoda, most probably titanosaurs, a group of large-bodied, , usually possessing a long neck and tail, with a small head.
The incomplete mandible attributed to A. wichmannianus is squared-off at the front with each dentary bone being ''L'' shaped. The teeth were restricted to the front of the lower jaw and were small and slender. The squared-off jaws suggest specialised feeding habits, such as feeding near a surface plane like low vegetation on the ground or floating plants in water. bones were, for the most part, not associated with each other but scattered throughout the formation. Consequently, many scientists believe that they may not all belong to the same type of animal. In particular, the very square lower jaw has frequently been suggested to belong to a rebbachisaurid sauropod similar to Nigersaurus. However, the jaw of Bonitasaura, described in 2004 , is similar in overall shape and is clearly associated with titanosaur skeletal remains, indicating that the lower jaw may belong to A. wichmannianus after all. In 2013 and 2018 respectively, Brasilotitan and Baalsaurus were described which also possessed squared-off jaws. It was noted that Brazilotitan, Bonitasaura, Antarctosaurus, and other titanosaurs show up three teeth per Dental alveolus (tooth socket) whereas the rebbachisaurid Nigersaurus shows up to seven teeth. Brazilotitan and Baalsaurus were described as a titanosaurs, closely related to A. wichmannianus.
The back of the skull and the remainder of the skeleton are usually regarded as titanosaurian by researchers, although they do not necessarily belong to the same type of titanosaur.
Powell compared the width of the cranium to the length of the limb bones of both A. wichmannianus and Saltasaurus; this led him to conclude that the skull was proportionally small in A. wichmannianus, this might imply that the skull and limb elements could belong to different individuals or a different Taxon. He noted, however, that the comparison was potentially misleading because the overall anatomy of Saltasaurus is shorter and stouter which might facilitate a bigger skull.
Von Huene assigned two tarsal (ankle) bones to A. wichmannianus, which he described as an astragalus and a calcanium. Powell suggested it's possible that the calcanium described by von Huene is actually the astragalus of a smaller individual. He also noted that the astragalus seems too small to belong to the same individual as the tibia, being only about half the width.
Von Huene described a Vertebra which was found close to the skull material. This vertebra was the first caudal, belonging to the base of the tail just after the sacrum (vertebrae attached to the hip). The vertebra features a biconvex centrum, a feature shared with other titanosaurs. Von Huene noted that the first caudal could possibly belong to Laplatasaurus .
With the exception of an incomplete cervical vertebra and the questionable first caudal, there are no vertebrae that link the skull to the limb material. There is a lack of field documentation to aid in the referral of all the material to one individual. Powell thought it was probable that von Huene correctly assigned the material to A. wichmannianus, arguing that von Huene would have been able to communicate with the discoverers and would have had access to photographs of the discovery site.
In 2016, using equations that estimate body mass based on the circumference of the humerus and femur of quadrupedal animals, it was estimated to be in weight. In 2019, Gregory S. Paul estimated the mass of A. giganteus in the range, based on newer titanosaur reconstructions. In 2020 Molina-Pérez and Larramendi estimated its length at and its weight similar to Paul's estimation at . Due to the incompleteness of the remains, any size estimates are subject to a large amount of error.
"Antarctosaurus" jaxarticus is known from a single femur which was briefly reported as resembling a femur attributed to Jainosaurus (then called "Antarctosaurus" septentrionalis). Paleontologist Teresa Maryańska noted that, whilst A. jaxarticus was named, it was not properly described or diagnosed. The femur possibly belongs to the titanosaur clade Lithostrotia.
The type specimen of "Antarctosaurus" brasiliensis is only known from three fragmentary bones that are titanosaurian in nature; a partial left femur GP-RD-2, a partial right humerus GP-RD-3, and an incomplete Vertebra (backbone) GP-RD-4. The femur is preserved and was estimated at if it were completed. The humerus is preserved and estimated at completed. The dorsal vertebra Vertebral body is long.
|
|