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The Alismatales ( alismatids) are an order of including about 4,500 species. Plants assigned to this order are mostly tropical or . Some grow in , some in . Perhaps the most important food crop in the order is the plant, Colocasia esculenta.


Description
The Alismatales comprise flowering plants of often aquatic and marshy habitats, and the only monocots known to have green embryos other than the . They also include the only marine angiosperms growing completely submerged, the . The are usually arranged in , and the mature seeds lack .

Both marine and freshwater forms include those with staminate flowers that detach from the parent plant and float to the surface. There they can pollinate carpellate flowers floating on the surface via long pedicels. In others, pollination occurs underwater, where pollen may form elongated strands, increasing chance of success. Most aquatic species have a totally submerged juvenile phase, and flowers are either floating or emerge above the water's surface. Vegetation may be totally submersed, have floating leaves, or protrude from the water. Collectively, they are commonly known as "water plantain".


Taxonomy
The Alismatales contain about 165 genera in 13 families, with a cosmopolitan distribution. , they are , diverging early in evolution relative to the and lineages. Together with the , the Alismatales are referred to informally as the alismatid monocots.


Early systems
The (1981) places the Alismatales in subclass , class = and includes only three families as shown: Cronquist's subclass Alismatidae conformed fairly closely to the order Alismatales as defined by APG, minus the Araceae.

The places the Alismatales in the superorder in the subclass = in the class = with the following families included:

In Tahktajan's classification (1997), the order Alismatales contains only the Alismataceae and Limnocharitaceae, making it equivalent to the Alismataceae as revised in APG-III. Other families included in the Alismatates as currently defined are here distributed among 10 additional orders, all of which are assigned, with the following exception, to the Subclass Alismatidae. Araceae in Tahktajan 1997 is assigned to the and placed in the Subclass Aridae; Tofieldiaceae to the and placed in the .


Angiosperm Phylogeny Group
The Angiosperm Phylogeny Group system () of 1998 and APG II (2003) assigned the Alismatales to the monocots, which may be thought of as an unranked clade containing the families listed below. The biggest departure from earlier systems (see below) is the inclusion of family Araceae. By its inclusion, the order has grown enormously in number of species. The family Araceae alone accounts for about a hundred genera, totaling over two thousand species. The rest of the families together contain only about five hundred species, many of which are in very small families.

The APG III system (2009) differs only in that the are combined with the Alismataceae; it was also suggested that the genus (of the ) could be separated into a monogeneric family, the , but the authors noted that more study was necessary before the Maundiaceae could be recognized.

In (2016), it was decided that evidence was sufficient to elevate Maundia to family level as the Maundiaceae. The authors considered including a number of the smaller orders within the Juncaginaceae, but an online survey of botanists and other users found little support for this "" approach. Consequently, the family structure for APG IV is:


Phylogeny
showing the orders of monocots ( Chase & Reveal) based on molecular phylogenetic evidence:


Further reading
  • B. C. J. du Mortier 1829. Analyse des Familles de Plantes : avec l'indication des principaux genres qui s'y rattachent. Imprimerie de J. Casterman, Tournay
  • W. S. Judd, C. S. Campbell, E. A. Kellogg, P. F. Stevens, M. J. Donoghue, 2002. Plant Systematics: A Phylogenetic Approach, 2nd edition. Sinauer Associates, Sunderland, Massachusetts .
  • (2025). 9781107012769, Cambridge University Press. .
  • , in


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