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Tonofibril
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Tonofibrils are cytoplasmic protein structures in tissues that converge at and . They consist of fine in epithelial cells that are anchored to the . They were discovered by Rudolf Heidenhain, and first described in detail by Louis-Antoine Ranvier in 1897.

Composition
Tonofilaments are intermediate filaments that make up tonofibrils in the epithelial tissue. In epithelial cells, tonofilaments loop through desmosomes. Electron microscopy has advanced now to illustrate the tonofilaments more clearly.

The protein is believed to be synthesized as a giant precursor protein, profilaggrin (>400 kDA in humans). When filaggrin binds to keratin intermediate filaments, the keratin aggregates into macrofibrils.

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