Product Code Database
Example Keywords: radiant silvergun -final $59
barcode-scavenger
   » » Wiki: Sexual Reproduction
Tag Wiki 'Sexual Reproduction'.
Tag

Sexual reproduction is a type of that involves a complex life cycle in which a (such as a or ) with a single set of () combines with another to produce an organism composed of cells with two sets of chromosomes ().John Maynard Smith & Eörz Szathmáry, The Major Transitions in Evolution, W. H. Freeman and Company, 1995, p 149 Sexual reproduction is the most common life cycle in multicellular , such as , and . Sexual reproduction does not occur in (organisms without ), but they have processes with similar effects such as bacterial conjugation, transformation and transduction, which may have been precursors to sexual reproduction in early eukaryotes.

In the production of sex cells in eukaryotes, diploid mother cells divide to produce haploid cells known as gametes in a process called that involves genetic recombination. The homologous chromosomes pair up so that their sequences are aligned with each other, and this is followed by exchange of genetic information between them. Two rounds of cell division then produce four haploid gametes, each with half the number of chromosomes from each parent cell, but with the genetic information in the parental chromosomes recombined. Two haploid gametes combine into one diploid cell known as a in a process called . The zygote incorporates genetic material from both gametes. Multiple , without change of the number of chromosomes, then form a multicellular diploid phase or generation.

In human reproduction, each cell contains 46 chromosomes in 23 pairs. Meiosis in the parents' gonads produces gametes that each contain only 23 chromosomes that are genetic recombinants of the DNA sequences contained in the parental chromosomes. When the nuclei of the gametes come together to form a fertilized egg or zygote, each cell of the resulting child will have 23 chromosomes from each parent, or 46 in total.

In plants only, the diploid phase, known as the , produces spores by meiosis that germinate and then divide by mitosis to form a haploid multicellular phase, the , that produces gametes directly by mitosis. This type of life cycle, involving alternation between two multicellular phases, the sexual haploid gametophyte and asexual diploid sporophyte, is known as alternation of generations.

The evolution of sexual reproduction is considered paradoxical, because asexual reproduction should be able to outperform it as every young organism created can bear its own young. This implies that an asexual population has an intrinsic capacity to grow more rapidly with each generation.John Maynard Smith The Evolution of Sex 1978. This 50% cost is a fitness disadvantage of sexual reproduction.Ridley M (2004) Evolution, 3rd edition. Blackwell Publishing, p. 314. The two-fold cost of sex includes this cost and the fact that any organism can only pass on 50% of its own genes to its offspring. One definite advantage of sexual reproduction is that it impedes the accumulation of genetic .

is a mode of natural selection in which some individuals out-reproduce others of a population because they are better at securing for sexual reproduction. It has been described as "a powerful evolutionary force that does not exist in asexual populations."


Evolution
The first evidence of sexual reproduction in eukaryotes is from the period, about 1 to 1.2 billion years ago.

Biologists studying propose several explanations for the development of sexual reproduction and its maintenance. These reasons include reducing the likelihood of the of deleterious mutations, increasing rate of adaptation to changing environments, dealing with competition, and masking deleterious mutations. All of these ideas about why sexual reproduction has been maintained are generally supported, but ultimately the size of the population determines if sexual reproduction is entirely beneficial. Larger appear to respond more quickly to some of the benefits obtained through sexual reproduction than do smaller population sizes.

Maintenance of sexual reproduction has been explained by theories that work at several levels of selection, though some of these models remain controversial. However, newer models presented in recent years suggest a basic advantage for sexual reproduction in slowly reproducing .

Sexual reproduction allows these species to exhibit characteristics that depend on the specific environment that they inhabit, and the particular survival strategies that they employ.


Sexual selection
In order to sexually reproduce, both males and females need to find a . Generally in animals is made by females while males compete to be chosen. This can lead to extreme efforts in order to reproduce, such as combat and display, or produce extreme features caused by a positive feedback known as a Fisherian runaway. Thus sexual reproduction, as a form of natural selection, has an effect on . Sexual dimorphism is where the basic vary between males and females of the same . Dimorphism is found in both and in secondary sex characteristics, body size, physical strength and morphology, biological ornamentation, and other bodily traits. However, sexual selection is only implied over an extended period of time leading to sexual dimorphism.Dimijian, G. G. (2005). Evolution of sexuality: biology and behavior. Proceedings (Baylor University. Medical Center), 18, 244–258.


Sex ratio
Apart from some , organisms which reproduce sexually have a 1:1 of male and female births. The English statistician and biologist outlined why this is so in what has come to be known as Fisher's principle. This essentially says the following:

# Suppose male births are less common than female.
# A newborn male then has better mating prospects than a newborn female, and therefore can expect to have more offspring.
# Therefore parents genetically disposed to produce males tend to have more than average numbers of grandchildren born to them.
# Therefore the genes for male-producing tendencies spread, and male births become more common.
# As the 1:1 sex ratio is approached, the advantage associated with producing males dies away.
# The same reasoning holds if females are substituted for males throughout. Therefore 1:1 is the equilibrium ratio.


Animals

Insects
Insect species make up more than two-thirds of all animal species. Most insect species reproduce sexually, though some species are facultatively . Many insects species have sexual dimorphism, while in others the sexes look nearly identical. Typically they have two sexes with males producing spermatozoa and females ova. The ova develop into eggs that have a covering called the , which forms before internal fertilization. Insects have very diverse mating and reproductive strategies most often resulting in the male depositing within the female, which she stores until she is ready for egg fertilization. After fertilization, and the formation of a zygote, and varying degrees of development, in many species the eggs are deposited outside the female; while in others, they develop further within the female and are born live.


Birds

Mammals
There are three extant kinds of mammals: , and , all with internal fertilization. In placental mammals, offspring are born as juveniles: complete animals with the present although not reproductively functional. After several months or years, depending on the species, the sex organs develop further to maturity and the animal becomes . Most female mammals are only during certain periods during their cycle, at which point they are ready to mate. Individual male and female mammals meet and carry out copulation. For most mammals, males and females .
(2020). 9780716740049, W.H. Freeman and Company. .
Research conducted by Patricia Adair Gowaty. Reported by


Fish
The vast majority of fish species lay eggs that are then fertilized by the male. BONY FISHES - Reproduction Some species lay their eggs on a substrate like a rock or on plants, while others scatter their eggs and the eggs are fertilized as they drift or sink in the water column.

Some fish species use internal fertilization and then disperse the developing eggs or give birth to live offspring. Fish that have live-bearing offspring include the and mollies or . Fishes that give birth to live young can be , where the eggs are fertilized within the female and the eggs simply hatch within the female body, or in , the male carries the developing young within a pouch, and gives birth to live young.

(2020). 9780761471943, Marshall Cavendish. .
Fishes can also be , where the female supplies nourishment to the internally growing offspring. Some fish are , where a single fish is both male and female and can produce eggs and sperm. In hermaphroditic fish, some are male and female at the same time while in other fish they are serially hermaphroditic; starting as one sex and changing to the other. In at least one hermaphroditic species, self-fertilization occurs when the eggs and sperm are released together. Internal self-fertilization may occur in some other species. One fish species does not reproduce by sexual reproduction but uses sex to produce offspring; is a unisex species that uses a form of called , where unfertilized eggs develop into embryos that produce female offspring. Poecilia formosa mate with males of other fish species that use internal fertilization, the sperm does not fertilize the eggs but stimulates the growth of the eggs which develops into embryos.


Reptiles

Amphibians

Mollusks

Plants
Animals have life cycles with a single diploid multicellular phase that produces haploid gametes directly by meiosis. Male gametes are called sperm, and female gametes are called eggs or ova. In animals, fertilization of the ovum by a sperm results in the formation of a diploid zygote that develops by repeated mitotic divisions into a diploid adult. Plants have two multicellular life-cycle phases, resulting in an alternation of generations. Plant zygotes germinate and divide repeatedly by mitosis to produce a diploid multicellular organism known as the sporophyte. The mature sporophyte produces haploid spores by meiosis that germinate and divide by mitosis to form a multicellular gametophyte phase that produces gametes at maturity. The gametophytes of different groups of plants vary in size. Mosses and other pteridophytic plants may have gametophytes consisting of several million cells, while angiosperms have as few as three cells in each pollen grain.


Flowering plants
are the dominant plant form on land and they reproduce either sexually or asexually. Often their most distinguishing feature is their reproductive organs, commonly called flowers. The produces which contain the male that produce sperm nuclei. For pollination to occur, pollen grains must attach to the stigma of the female reproductive structure (), where the female gametophytes (ovules) are located inside the . After the pollen tube grows through the carpel's style, the cell nuclei from the pollen grain migrate into the ovule to fertilize the egg cell and endosperm nuclei within the female gametophyte in a process termed double fertilization. The resulting zygote develops into an embryo, while the triploid endosperm (one sperm cell plus two female cells) and female tissues of the ovule give rise to the surrounding tissues in the developing seed. The ovary, which produced the female gametophyte(s), then grows into a , which surrounds the seed(s). Plants may either or .

In 2013, flowers dating from the (100 million years before present) were found encased in amber, the oldest evidence of sexual reproduction in a flowering plant. Microscopic images showed tubes growing out of pollen and penetrating the flower's stigma. The pollen was sticky, suggesting it was carried by insects.

Nonflowering plants like , and use other means of sexual reproduction.


Ferns
Ferns mostly produce large diploid with , roots and leaves. Fertile leaves produce that contain haploid . The spores are released and germinate to produce short, thin gametophytes that are typically heart shaped, small and green in color. The gametophyte , produce both motile sperm in the and egg cells in on the same or different plants. After rains or when dew deposits a film of water, the motile sperm are splashed away from the antheridia, which are normally produced on the top side of the thallus, and swim in the film of water to the archegonia where they fertilize the egg. To promote out crossing or cross fertilization the sperm are released before the eggs are receptive of the sperm, making it more likely that the sperm will fertilize the eggs of different thallus. After fertilization, a is formed which grows into a new sporophytic plant. The condition of having separate sporophyte and gametophyte plants is called alternation of generations. Other plants with similar life cycles include , , and .


Bryophytes
The , which include , and , reproduce both sexually and vegetatively. They are small plants found growing in moist locations and like ferns, have motile sperm with and need water to facilitate sexual reproduction. These plants start as a haploid spore that grows into the dominant gametophyte form, which is a multicellular haploid body with leaf-like structures that . Haploid gametes are produced in antheridia (male) and archegonia (female) by mitosis. The sperm released from the antheridia respond to chemicals released by ripe archegonia and swim to them in a film of water and fertilize the egg cells thus producing a zygote. The divides by mitotic division and grows into a multicellular, diploid sporophyte. The sporophyte produces spore capsules (), which are connected by stalks () to the archegonia. The spore capsules produce spores by meiosis and when ripe the capsules burst open to release the spores. Bryophytes show considerable variation in their reproductive structures and the above is a basic outline. Also in some species each plant is one sex (dioicous) while other species produce both sexes on the same plant (monoicous).
(1988). 9780195063943, Oxford University Press.


Fungi
are classified by the methods of sexual reproduction they employ. The outcome of sexual reproduction most often is the production of that are used to survive inclement times and to spread. There are typically three phases in the sexual reproduction of fungi: , and . The cytoplasm of two parent cells fuse during plasmogamy and the nuclei fuse during karyogamy. New haploid gametes are formed during meiosis and develop into spores. The adaptive basis for the maintenance of sexual reproduction in the and () was reviewed by Wallen and Perlin. They concluded that the most plausible reason for maintaining this capability is the benefit of , caused by a variety of stresses, through recombination that occurs during .


Bacteria and archaea
Three distinct processes in are regarded as similar to eukaryotic sex: bacterial transformation, which involves the incorporation of foreign DNA into the bacterial chromosome; bacterial conjugation, which is a transfer of DNA between bacteria, but the plasmids are rarely incorporated into the bacterial chromosome; and gene transfer and genetic exchange in archaea.

Bacterial transformation involves the recombination of genetic material and its function is mainly associated with DNA repair. Bacterial transformation is a complex process encoded by numerous bacterial genes, and is a bacterial adaptation for DNA transfer. This process occurs naturally in at least 40 bacterial species. For a bacterium to bind, take up, and recombine exogenous DNA into its chromosome, it must enter a special physiological state referred to as competence (see Natural competence). Sexual reproduction in early single-celled eukaryotes may have evolved from bacterial transformation,Bernstein H, Bernstein C, Michod RE. (2012) " DNA Repair as the Primary Adaptive Function of Sex in Bacteria and Eukaryotes". Chapter 1, pp. 1–50, in DNA Repair: New Research, Editors S. Kimura and Shimizu S. Nova Sci. Publ., Hauppauge, New York. Open access for reading only. or from a similar process in (see below).

On the other hand, bacterial conjugation is a type of direct transfer of DNA between two bacteria mediated by an external appendage called the conjugation pilus. Bacterial conjugation is controlled by that are adapted for spreading copies of the plasmid between bacteria. The infrequent integration of a plasmid into a host bacterial chromosome, and the subsequent transfer of a part of the host chromosome to another cell do not appear to be bacterial adaptations.

(2020). 9780763766320, Jones and Bartlett Publishers. .

Exposure of hyperthermophilic archaeal Sulfolobus species to DNA damaging conditions induces cellular aggregation accompanied by high frequency exchange. Ajon et al. hypothesized that this cellular aggregation enhances species-specific DNA repair by homologous recombination. DNA transfer in Sulfolobus may be an early form of sexual interaction similar to the more well-studied bacterial transformation systems that also involve species-specific DNA transfer leading to homologous recombinational repair of DNA damage.


See also


Further reading


External links

Page 1 of 1
1
Page 1 of 1
1

Account

Social:
Pages:  ..   .. 
Items:  .. 

Navigation

General: Atom Feed Atom Feed  .. 
Help:  ..   .. 
Category:  ..   .. 
Media:  ..   .. 
Posts:  ..   ..   .. 

Statistics

Page:  .. 
Summary:  .. 
1 Tags
10/10 Page Rank
5 Page Refs
2s Time