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Sander (fish)
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Sander (formerly known as Stizostedion) is a of predatory in the family , which also includes the , , and . They are also known as "pike-perch" because of their resemblance to fish in the unrelated (pike) family. They are the only genus in the tribe Luciopercini, which is one of two tribes in the .

The earliest known fossils of this genus are partial jaw and vertebrae elements from the middle ()-aged Wood Mountain Formation of , Canada.


Characteristics
Sander species have elongated and laterally compressed bodies and they range in from in the Volga pikeperch ( S. volgensis) to in the zander ( S. lucioperca). The species within the genus share canine-like teeth that are at their largest in the zander, and although they are not present in adult Volga pikeperches, they do possess them as juveniles. in addition, they have thin rows of teeth on their jaws, vomer, and palatines, the preopercle shows strong serrations along its edges, a continuous reaches all the way from the head to the , and this is flanked by additional lateral lines, one each on the upper and lower lobes of the deeply forked . Further features in common include the absence of genital papillae, seven or eight branchiostegal rays, 12–13 soft rays in the , and the eye has a reflective layer behind the , known as a , which is an adaptation for seeing in low-light conditions. The species in the genus Sander are largely as adults.


Species
The genus includes these species: The fossil species Sander svetovidovi Kovalchuk, 2015 is known from the Late Miocene of Ukraine.


Phylogeny
Phylogenetic relationships of the species of genus Sander based on the concatenated data set of six gene regions and a Bayesian analysis. Romanichthys valsanicola is the nearest living relative of the genus Sander and is used as an outgroup to root the tree.

This is not universally accepted, though, and the ( Romanichthys valsanicola) has been more recently placed within the genus .

(2025). 9789401772273, Springer, Dordrecht.

Two are within the genus, a Eurasian one and a North American one, which separated from a common ancestor around 20.8 million years ago (Mya) in the , when the North Atlantic Land Bridge connecting Europe to eastern North America subsided. The Eurasian clade then speciated from 13.8 Mya, while the two North American species speciated around 5.4 Mya.

The relatively old divergences given for North American and Eurasian Sander are supported by the discovery of a fossil Sander from the (16.3 to 13.6 mya)-aged Wood Mountain Formation of Canada. Prior to this discovery, it was suggested that Sander may have potentially been a much more recent immigrant to North America, potentially as young as the or even . In Eurasia, fossils of the extant S. lucioperca and the extinct S. svetovidovi are known from the and early Pliocene of Ukraine. These fossils also suggest a coexistence between Sander and their close relative, the now-extinct . It has been theorized that Sander was more tolerant than Leobergia to the global cooling that occurred during the Pliocene, leading to the extinction of the latter.


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