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Mesonychidae (meaning "middle ") is an extinct family of small to large-sized - . They were to and during the to the , and were the earliest group of large carnivorous mammals in . Once considered a to , recent evidence now suggests no close connection to any living mammal. Mesonychid has long been disputed and they have captured popular imagination as " on ", that combine features of both and . and have similar features to early , and the family was long thought to be the ancestors of . Recent discoveries have overturned this ; the consensus is that whales are highly derived artiodactyls. Some researchers now consider the family a sister group either to whales or to artiodactyls, close relatives rather than direct ancestors. Other studies define as basal to all ungulates, occupying a position between and . In this case, the resemblances to early whales would be due to convergent evolution among ungulate-like that developed adaptations related to hunting or eating meat.


Description
The mesonychids were an unusual group of with a specialized dentition featuring tri-cuspid upper molars and high-crowned lower molars with shearing surfaces. They had large heads with relatively long necks. Over time, the family evolved foot and leg adaptations for faster running, and jaw adaptations for greater bite force. Like the Paleocene family , mesonychids were once viewed as primitive , and the diet of most genera probably included meat or fish. Various genera and species coexisted in some locations, as hunters and omnivores or scavengers. In contrast to arctocyonids, the mesonychids had only four digits furnished with hooves supported by narrow fissured end phalanges.


Evolutionary history
They first appeared in the Early , undergoing numerous speciation events during the Paleocene, and . Mesonychids fared very poorly at the close of the Eocene epoch, with only one genus, , surviving into the Early epoch.

Mesonychids probably originated in , where the most primitive mesonychid, , is known from the early Paleocene. They were also most diverse in Asia where they occur in all major Paleocene . Since other such as the and were either rare or absent in these animal communities, mesonychids most likely dominated the large in the Paleocene of Asia. Throughout the Paleocene and Eocene, several genera, including , and would radiate out from their ancestral home in Asia and into Europe and North America, where they would give rise to new mesonychid genera. These animals would have migrated to North America via the Bering land bridge.


Taxonomy
Mesonychidae was named by Cope (1880). Its type genus is Mesonyx. It was assigned to by Cope (1880); to Creodonta by Cope (1889); to by Peterson (1919); to Mesonychia by Carroll (1988) and Zhou et al. (1995); and to Cete by Archibald (1998);J. D. Archibald. 1998. Archaic ungulates ("Condylarthra"). In C. M. Janis, K. M. Scott, and L. L. Jacobs (eds.), Evolution of Tertiary Mammals of North America 1:292-331 and to Mesonychia by Carroll (1988), Zhou et al. (1995), Geisler and McKenna (2007) and Spaulding et al. (2009).. Retrieved 11 August 2013.


Classification
Family Mesonychidae


Notes

Cited sources

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