Characidae, the characids, is a family of Fresh water subtropical and tropical fish belonging to the order Characiformes. They are found throughout much of Central America and South America, including such major waterways as the Amazon River and Orinoco Rivers.[Nelson (2006)] These fish vary in length; many are less than .
The name "Characin" is a historical one,[Characinae, recently narrowly defined, covers only twelve genera and 79 species closely related to Charax (George M.T. Mattox, Monica Toledo-Piza, "Phylogenetic study of the Characinae (Teleostei: Characiformes: Characidae)" Zoological Journal of the Linnean Society, 165.4:809–915, August 2012).] but scientists today tend to prefer "characids" to reflect their status as a, by and large, monophyletic group (at family rank). This family includes some of the first characiforms to be described to science, such as Charax and Tetragonopterus, and thus lend their name to the order, as well as to common names such as "characin" and "tetra".
Past taxonomic treatments had a much more expansive definition of the family, including numerous South American fish families such as the Piranha and Bryconidae, as well as the African Alestidae. Following multiple taxonomic revisions, this was eventually restricted to just the American "tetra" type characins by the 2010s. However, even this definition of Characidae was found to obscure much of the evolutionary diversity within the group, and in 2024 the "tetra" families Acestrorhamphidae and Stevardiidae were split out of the Characidae, leaving it with just five subfamilies.
Classification
Taxonomy
The following classification is based on
Eschmeyer's Catalog of Fishes (2025):
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Family Characidae
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Subfamily Aphyocharacinae C. H. Eigenmann, 1909
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Amazonichthys Esguícero & Mendonça, 2023
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Aphyocharacidium Géry, 1960
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Aphyocharax Günther, 1868
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Cyanogaster Mattox, Ralf Britz, Toledo-Piza & Marinho, 2013
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Leptagoniates Boulenger, 1887
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Paragoniates Steindachner, 1876
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Phenagoniates Eigenmann & C. B. Wilson, 1914
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Prionobrama Fowler, 1913
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Xenagoniates Myers, 1942
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Subfamily Cheirodontinae C. H. Eigenmann, 1915
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Acinocheirodon Malabarba & Weitzman, 1999
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Amblystilbe Fowler, 1940
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Aphyocheirodon C. H. Eigenmann, 1915
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Cheirodon Girard, 1855
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Cheirodontops Schultz, 1944
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Compsura C. H. Eigenmann, 1915
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Ctenocheirodon Malabarba & Jerep, 2012
[Malabarba, L.R. & Jerep, F.C. (2012): A New Genus and Species of Cheirodontine Fish from South America (Teleostei: Characidae). Copeia, 2012 (2): 243–250.]
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Heterocheirodon Malabarba, 1998
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Holoshesthes C. H. Eigenmann, 1903
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Kolpotocheirodon Malabarba & Weitzman, 2000
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Macropsobrycon C. H. Eigenmann, 1915
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Nanocheirodon Malabarba, 1998
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Odontostilbe Cope, 1870
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Prodontocharax C. H. Eigenmann & Pearson, 1924
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Protocheirodon Vari, Melo & Oliveira, 2016
[Vari, R.P., Melo, B.F. & Oliveira, C. (2016): Protocheirodon, a new genus of Characidae (Teleostei: Characiformes) with the redescription of the poorly known Protocheirodon pi. Neotropical Ichthyology, 14 (2): e150154.]
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Pseudocheirodon Meek & Hildebrand, 1916
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Saccoderma Schultz, 1944
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Serrapinnus Malabarba 1998
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Subfamily Exodontinae Fowler, 1958
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Subfamily Tetragonopterinae Theodore Gill, 1858
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Subfamily Characinae Latreille, 1825
Former members
This family has undergone a large amount of systematic and taxonomic change. More recent revision has moved many former members of the family into their own related but distinct families – the of the genus Nannostomus are a typical example, having now been moved into the Lebiasinidae, the assorted predatory species belonging to Hoplias and Hoplerythrinus have now been moved into the Erythrinidae, and the sabre-toothed fishes of the genus Hydrolycus have been moved into the Cynodontidae. The former subfamily Alestiinae was promoted to family level (Alestiidae) and the subfamilies Crenuchinae and Characidiinae were moved to the family Crenuchidae.
Other fish families that were formerly classified as members of the Characidae, but which were moved into separate families of their own during recent taxonomic revisions (after 1994) include Acestrorhynchidae, Anostomidae, Chilodidae, Citharinidae, Ctenoluciidae, Curimatidae, Distichodontidae, Gasteropelecidae, Hemiodontidae, Hepsetidae, Parodontidae, Prochilodontidae, Serrasalmidae, and Triportheidae.[Oliveira, C., Avelino, G.S., Abe, K.T., Mariguela, T.C., Benine, R.C., Orti, G., Vari, R.P., & Correa e Castro, R.M. (2011): Phylogenetic relationships within the speciose family Characidae (Teleostei: Ostariophysi: Characiformes) based on multilocus analysis and extensive ingroup sampling. BMC Evolutionary Biology, 11: 275. doi: 10.1186/1471-2148-11-275] In 2024, the families Stevardiidae and Acestrorhamphidae, containing a high proportion of the famous ornamental aquarium tetras, were also split out of the family, in addition to the small family Spintherobolidae.
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de Lucena, Carlos Alberto Santos (2003): New characid fish, Hyphessobrycon scutulatus, from the rio Teles Pires drainage, upper rio Tapajós system (Ostariophysi: Characiformes: Characidae). Neotropical Ichthyology 1(2): 93–96. PDF fulltext
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Géry, Jacques (1977): Characoids of the World.
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Nelson, Joseph S. (2006): Fishes of the World. John Wiley & Sons, Inc.